Redescription of the genus Allocryptobia Viette, 1951 (Lepidoptera: Cossidae)

Based on the study of male and female genitalia and the external features of Cryptobia musae Herrich-Schäffer, [1854], the type species of the genus Allocryptobia Viette, 1951, this genus is redescribed. Images of the type specimens, male and female genitalia, a new generic diagnosis, and a distribution map are given. Lectotypes of Cryptobia musae Herrich-Schäffer, [1854] and Cossus mucoreus Herrich-Schäffer, [1853] are designated. Allocryptobia musae is recorded for the fauna of Honduras, Panama and Colombia for the first time.


Introduction
The Cossid fauna of Neotropics is poorly known.Only the subfamily Cossulinae (DAVIS et al., 2008) has been relatively well treated.We have initiated a systematic revision of the South American carpenter-moths with redescribing of poorly known genera (YAKOVLEV, 2014;PENCO et al., 2016;YAKOVLEV et al., 2016).Additionally, we have recently published a preliminary list of the Cossidae (PENCO & YAKOVLEV, 2015) of Argentina, and descriptions of several new species (PENCO & YAKOVLEV, 2017;YAKOVLEV et al., 2017).
It is known that the type material and other collections of Herrich-Schäffer have been deposited in various European museums (HÄUSER et al., 2003).A part of this material from the collection of Kaden, which served as a basis for iconography by HERRICH-SCHÄFFER (1850-1858), was recently found in the entomological collection of ZISP; it includes several types of Cossidae from South America.

Material and methods
Male and female genitalia were mounted in micro tubes (under specimens on a pin) and examined with a Nikon SMZ 800 n microscope.Images of genitalia were taken with the Olympus XC 50 camera.Images of imago were taken by the digital camera of Apple iPhone 7, illuminated in Lightbox.The images were processed using CorelDraw software.

Results
We examined syntypes of Cryptobia musae (male and female) and Cossus mucoreus (female) kept in the collection of ZISP, as well as some new material of the first species, collected in Panama and Colombia.Because Figure 165 in HERRICH-SCHÄFFER (1850-1858) depicts the female of C. musae, we designate the female specimen as a lectotype and the male one as the paralectotype (article 74B of ICZN, 1999).Figure 39 in HERRICH-SCHÄFFER (1850-1858) for C. mucoreus also illustrates a female, and as there is no evidence that it is a unique specimen, we designate it as a lectotype.

Redescription of the genus is given below based on its type species.
Male: Antenna short, bipectinate in proximal half, filiform in distal half.Fore wing relatively narrow, apically rounded, with specific black pattern.Hind wing short, relatively wide, almost completely black basally, with dense reticulated black pattern distally.
Female: Antenna short, filiform.Wings significantly wider than those of male.Male genitalia (Fig. 9): Uncus long, triangle, with beak-shaped point on top; gnathos arm short, thick, fused with vinculum by membrane; valve relatively narrow, costal edge slightly curved in medium third, abdominal edge strongly curved in proximal third, small lanceolate harpe in medium third of abdominal edge, apex semicircular; juxta small, with long leaf-like lateral processes, strongly fused with phallus; saccus tiny, semicircular; phallus thick, shorter than valve, slightly curved in medium third, robust cornutus in lateral surface of vesica.
Female genitalia (Fig. 10): Ovipositor very long.Ostium poorly immersed, cup-like; ductus thick, of medium length, with sack-shaped bulla extending from it on a long membrane duct; bursa big, oval, with small stellate signum on lateral surface, ductus seminalis thin, extending from top of bursa; anterior apophyses much longer than posterior ones; transverse oblique incisions on lateral surface of ovipositor; papillae anales semicircular.Distribution.Brazil and Venezuela.

Discussion
Basing on the male genital characters of the type species, the genus Allocryptobia belongs to the group of genera of the subfamily Zeuzerinae having one harpe on the lower edge of valve.Now this group includes the South American genera Brypoctia Schoorl, 1990 (type species Xyleutes strigifer Dyar, 1910), Morpheis Hübner, [1820] (type species Phalaena pyracmon Cramer, 1782) and some Australian genera combined by the KALLIES & HILTON (2012) into the "Sympycnodes digitata group".We are stated that apomorphies for the genus Allocryptobia include the specific habitus and the tiny sacculus, the short robust phallus and the gnathos arms fused with vinculum in the male genitalia.
A number of questions on the taxonomy of the genus Allocryptobia remain controversial.The most important is the synonymy of C. mucoreus with C. phobifera established by SCHOORL (1990: 164) with the following argument: "The holotype of phobifera is similar to a male specimen from Venezuela in the BMNH, which has been identified together with a female from the same locality as belonging to mucorea.Dyar notes that the hind wings are partly without scales due to damage.It appears to be a characteristic that a moderately broad band along the hind wing dorsum is unsealed, since the male in the BMNH has this too.There are apparently no good differences between the two species.Therefore, phobifera is synonymized with mucorea."However, the conspecificity of the male and female of C. mucoreus from Venezuela (BMNH) and the male from Paraguay (holotype Cryptobia phobifera) is doubtful (Figures 7 and 8, respectively).
Additionally, SCHOORL (1990) mentioned the pronounced sexual dimorphism as the most important characteristic of the genus Allocryptobia.This statement is based on comparing the external features of both sexes of Cryptobia mucoreus, however, in the females and males of the type species sexual dimorphism is only slightly expressed.We provisionally leave C. mucoreus in the genus Allocryptobia but its taxonomic position, as well as conspecifity with C. phobifera, needs further study.