Tortricidae from Ethiopia, 4 (Insecta: Lepidoptera)

Forty species of Tortricidae from North-Western Ethiopia are recorded of which fifteen species are described as new ( Eugnosta amharana Razowski & Trematerra, sp. n.; Procrica sinuata Razowski & Trematerra, sp. n.; Metamesia flava Razowski & Trematerra, sp. n.; Clepsis paragongyla Razowski & Trematerra, sp. n.; Epichoristodes fekensae Razowski


Introduction
Tortricidae of Ethiopia are still very little known and since the MEYRICK (1932) publication of the results of an expedition to Abyssinia (Ethiopia) in the years 1926-1927 only the collecting trips discussed below were made. The history of the entomological expeditions of the University of Molise steady human and livestock pressures which is believed to reduce the area of the Wof-Washa Forest as time goes on. Some of the common woody species in Wof-Washa Forest include Juniperus procera Hochst. ex Endl, Afrocarpus falcatus Thunberg, Polyscias fulva (Hiern) Harms, Hypericum revolutum Vahl, Ekebergia capensis Sparrm., Bersama abyssinica Fresen., Olea hochstetteri Bak., Celtis africana Burm., Hagenia abyssinica (Bruce) Gmel., and Prunus africana (Hook.) Kalkm. The collections site is at 2900 m/a.s.l. (BEKELE, 1994;TEKETAY & BEKELE, 1995).

Material and methods
Adults were collected during the night using UV-LED light traps and from a white sheet placed behind the same light. Genitalia were prepared using standard methods, the abdomen was macerated in 10% KOH and dissected under a stereoscopic microscope, the genitalia were separated and mounted in euparal on a glass slide. Adults and slides are housed in P. Trematerra Collection, Campobasso (Italy). Description: Male, wing span 18 mm. Head and thorax yellow. Forewing almost uniformly broad; costa weakly convex; termen straight, slightly oblique. Ground colour yellow, basal fourth of costa and base of dorsum suffused brown, remaining part of latter spotted brown, suffused brownish-grey, browner at tornus; concolous darker suffusion from before tornus to costal half of median fascia. Markings brown extending from mid-dorsum to beyond mid-costa, oblique from median cell to latter. Cilia damaged, remnants yellowish. Hindwing cream distinctly strigulated greyish brown; cilia damaged.
Female unknown. Dignosis: E. amharana is related to E. marginana Aarvik, 2010 from Uganda but the socius of latter is short and the median part of transtilla without terminal incision.
Female unknown. Diagnosis: P. sinuata is related to P. parisii Razowski & Trematerra, 2010 from Bale Mountains, RAZOWSKI & TREMATERRA (2010) illustrated this species but the male genitalia were originally mistakenly numbered (RAZOWSKI et al., 2018). From P. parisii this species differs in colouration and the shape of the uncus.
Female unknown. Diagnosis: M. flava, in facies is similar to M. incepta (Meyrick, 1912) but differs from it by a lack of black forewing dots on a dark yellow ground colour. The male genitalia are similar to those of M. designata (Meyrick, 1912)  Description: Male, wing span 16 mm. Head orange yellow, labial palpus orange yellow; thorax brown. Forewing not expanding terminad; costa weakly convex; termen slightly oblique, tolerably straight. Ground colour brownish orange strongly suffused rust brown, yellow along termen. Markings dark rust brown, diffuse, consisting of median fascia and subapical blotch. Cilia orange yellow. Hindwing brownish grey, in apical part yellow; cilia grey and yellowish, respectively.
Female similar to male with paler, yellower ground colour covering broad terminal area; the marking are more distinct than in the male. Hindwing grey-brown, yellowish apical area weak.
Diagnosis: C. paragongyla is related to C. gongyla Razowski, 2014 from Congo but is distinct by having the slender markings, broad uncus, short aedeagus, short processes of transtilla, and short arms of vinculum.
Etymology: The name refers to similarity with C. gongyla Razowski. Description: Male, wing span 19 mm. Head and thorax yellow brownish; labial palpus brownish. Forewing not expanding terminad; costa almost uniformly broad; termen straight, oblique. Wing yellow, basal part of costa brown. Numerous black dots over the wing except for costa and dorsum, agglomerate at costal half of termen. Cilia slightly paler than wing. Hindwing grey, yellowish in apical part; cilia yellowish.

Epichoristodes fekensae
Variation: Paratype forewing weakly dotted; larger spot at end of median cell. Female, wing span 20 mm. Similar to male with black dots over the wing except for costa and basal part, agglomerate at subterminal bloth.
Diagnosis: E. fekensae is similar to E. imbriculata (Meyrick, 1936) from Zaire but E. fekensae has a weakly convex base of the forewing costa and slenderer, subterminally thorny aedeagus; from E. intensa (Meyrick, 1921), from Natal, South Africa this species differs also by the shape of the aedeagus, sacculus and uncus.
Diagnosis: Thiopeia is related and in male genitalia similar to Choristoneura Lederer, 1859 and Mabilleodes Diakonoff, 1960, but differs from the former by a very small caulis, the setose sacculus, presence of sterigma lobes, broad ductus bursae, absence of cestum, minute blade of signum and large, strongly sclerotized subgenital sternite. It is also similar to the Madagascan Cornusaccula Diakonoff, 1968 but this genus haracterizes by short sacculus armed with long setae.
Etymology: The generic name is an anagram of the country name Ethiopia, from which the typespecies of the genus comes. Description: Male, wing span 19 mm. Head yellow cream, labial palpus yellow cream; thorax brownish. Forewing not expanding terminad, broadest submedially; costa convex basally; apex pointed; termen straight, slightly oblique. Ground colour yellow cream delicately strigulated brownish, mixed brown along costa and dorsum, slightly so terminally. Markings brownish with brown dots or strigulae: basal blotch ill-defined, median fascia interrupted near middle, atrophying at dorsum, with dark brown blotch before dorsum; subapical blotch elongate triangular reaching apex of wing; elongate mark at vein M 2 subterminally; cilia yellow. Hindwing greyish brown, yellowish at apex where sparsely spotted brownish; cilia yellowish.
Female, wing span 15-22 mm, similar to male but forewing ground colour much paler, venation in tornal area brownish, basal blotch ill-defined; median fascia brownish yellow, twice interrupted; brownish grey area of hindwing more or less reduced. In one example forewing markings are weakly developed while suffusions dark, distinct.
Male genitalia ( Figure 40) and female genitalia ( Figure 41) as described with the genus. Diagnosis: T. chokeana is the only representative of the genus; it can be compared with Tortrix diametrica Meyrick, 1932 from Madagascar but the latter has a broad valva and a thorn from the end of the sacculus. From Cornusaccula periopa Diakonoff, 1960 this species differs chiefly by having a short sacculus and a simple aedeagus.
Etymology: The name refers to Choke territory where the species was collected. Description: Female, wing span 17 mm. The specimen is damaged thus the wing markings are impossible to be described accurately.
Diagnosis: T. africana is distinct by its genitalia; in male genitalia it resembles the Palaearctic T. irinae Budashkin, 1990 but T. irinae has small, slender uncus and subdorsal row of spines in the disc of the valva. In female genitalia, the sterigma of T. africana resembles that of another Palaearctic species, T. couleruana (Duponchel, 1834) but the latter has an elongate, telescopic ovipositor and a rudimentary signum and, as T. irinae, white forewing ground colour.
Etymology: The name refers to Africa. Description: Wing span 18 mm. Head and thorax brown. Forewing slightly expanding terminally; costa weakly, uniformly broad throughout; termen sinuate beneath apex. Ground colour cream brown in form of slender interfascia, suffused brown. Markings broad, diffuse, dark brown; costa and termen concolorous. White line from beneath apex to mid-termen. Cilia brown. Hindwing greyish brown, cilia brownish cream.
Etymology: The name refers to the locality of Wof-Washa were the new species was collected. Description: Male, wing span 15 mm. Head yellow-brown, also palpus yellow-brown; thorax brown. Forewing not expanding terminad; costa slightly convex; termen moderately oblique, concave beneath apex. Ground colour brownish yellow with browner suffusions and dots; costal strigulae small, yellowish cream, divisions brown. Markings brown: basal blotch subdivided into diffuse parts with some dark brown strigulae and spots; median fascia marked brown medially; subapical blotch brown, also with brown marks. Cilia damaged, remnant brown and yellowish. Hindwing brown, paler basally; cilia similar.
Female unknown. Diagnosis: M. valvalobata is related to the South African (Natal) M. calliarma (Meyrick, 1909) and M. nereidopa (Meyrick, 1927) from Kenya differing from them chiefly in the shape of the cucullus.
Diagnosis: C. triangularis is similar and closely related to the South African C. tumulata (Meyrick, 1908) but differs from it in having a very short, subtriangular uncus, strongly curved, long socii, a convex caudal edge of the cucullus, and a long, slender valva. In C. tumulata the most importand differing character is a slender uncus. The female of this species has a longer sclerite antrum and a shorter subgenital sternite than C. tumulata.
Female unknown. Diagnosis: In facies, T. parimitans is similar to Cryptoschesis imitans Diakonoff, 1988 from Madagascar but differs from it chiefly in having a short, broad aedeagus.
Etymology: The species is named after C. imitans. Description: Male, wing span 14 mm. Head and proximal part of thorax orange cream, front orange; remaining parts of thorax brownish, palpi orange cream. Forewing expanding terminad; termen straight, moderately oblique. Ground colour creamish more or less suffused brownish and ferruginous, strigulated and reticulated brown. Markings: dorsobasal blotch edged rust brown followed by brown suffusion; median fascia rudimentary in costal part brownish; subterminal fascia indistinct parallelly edged, reaching posterior part of termen. Cilia brownish cream. Hindwing grey cream with brownish grey suffusions and similar cilia.
Female unknown. Diagnosis: In facies, T. machakelana is similar to T. batrachopa but differs from it by having an ill-defined subterminal fascia; in genitalia, it resembles T. leucotreta Meyrick, 1913 from which it differs chiefly by longer cucullus and ventroterminal part of the aedeagus. T. machakelana differs from the two mentioned species chiefly by having a thorn-like process of the neck of valva.
Etymology: The name refers to the territory of Machakel Woreda where the new species was collected.