New and interesting Portuguese Lepidoptera records from 2019 (Insecta: Lepidoptera)

Novos e interessantes registos portugueses de Lepidoptera em 2019 (Insecta: Lepidoptera)

Nuevas e interesantes citas portuguesas de Lepidoptera en 2019 (Insecta: Lepidoptera)

Introduction

This paper is the fourteenth in the series of annual summaries of new knowledge of Portuguese Lepidoptera. It gives records of species of Lepidoptera added to the Portuguese fauna in 2019, together with new province records not included in the checklist (CORLEY, 2015). Additional data include new data on larval food-plants within the country and second and third records of species for the country, which are only indicated when they are not in new provinces. Papers published in 2019 and part of 2020 that relate to the Portuguese Lepidoptera fauna are listed and briefly summarised. Finally, an Appendix lists the new species for Portugal separately, with numbers indicating their position in the checklist; new genera for Portugal have author and year of publication given.

Fourteen species new for Portugal are listed below, of which three are new for the Iberian Peninsula. Two species are removed from the Portuguese list. A few of the new species listed here have been previously listed for Portugal, but the records were rejected in CORLEY (2015), as being erroneous or unsubstantiated.

In CORLEY et al. (2019) the number of Lepidoptera species recognised from Portugal was 2709. With the current paper and other papers mentioned herein, this total has risen to 2723.

Material and Methods

Most species were captured at light in traps of various kinds, or over or beside a white sheet. For specimens not taken at light, the means of capture is given. Specimens are retained in the collections of the original recorders, unless otherwise stated. However, a few records are based only on photographic evidence.

The number of new district records in 2019 is unusually high, partly due to fieldwork carried out in the relatively poorly worked districts of Baixo Alentejo and Ribatejo, but also to records received from the project Invertebrates Red List of Continental Portugal. This involved a series of fieldwork sessions from March to August, each several days long, moving to a new locality each day, with four workers from a team of thirteen in the field in each session. In the list of records below, records from this source have the recorder given as “Tagis”. The full Tagis team is given below following Recorders.

The order and nomenclature of families and species has been revised in accordance with the Portuguese list (CORLEY, 2015). The nomenclature of plant names follows the EURO+MED PLANT- BASE where possible.

The entry for species new for Portugal concludes with a summary of the known European distribution, and available information on the larval food-plant, given in square brackets if the information comes from outside Portugal.

Localities with UTM squares and altitude: (District in brackets)

Recorders and Determiners

1. Wilfried Arnscheid

2. Érica Brunheta

3. Martin Corley

4. Agostinho Fernandes

5. Isaías Ferreira

6. Sónia Ferreira

7. Dieter Fritsch

8. Darinka Gonzalez

9. David Grundy

10. Edmundo Jesus

11. André Lameirinhas

12. Eduardo Marabuto

13. Teodoro Monteiro

14. João Nunes

15. Pedro Pires

16. Jorge Rosete

17. Carlos Silva

18. Peder Skou

19. Rachel Terry

20. Martin Townsend

21. Ana Valadares

22. José Luis Yela García

Tagis team:

1. Sandra Antunes

2. Francisco Barros

3. Gonçalo Costa

4. Ricardo Costa

5. Rui Félix

6. Patrícia Garcia-Pereira

7. Ana Rita Gonçalves

8. Sam Martins

9. Tatiana Moreira

10. Sílvia Pina

11. Ricardo Ramirez

12. Renata Santos

13. Albano Soares

Abbreviations and symbols

coll. collection

comm communicated by

conf confirmed by

det. determined by

SMNK Staatliches Museum für Naturkunde, Karlsruhe, Germany

* New for Portugal, i.e. not listed for Portugal in CORLEY (2015).

** New for the Iberian Peninsula.

Provinces:

AAL Alto Alentejo

ALG Algarve

BA Beira Alta

BAL Baixo Alentejo

BB Beira Baixa

BL Beira Litoral

DL Douro Litoral

E Estremadura

M Minho

R Ribatejo

TM Trás-os-Montes

List of families and species

MICROPTERIGIDAE

Micropterix herminiella Corley, 2007

E: Vale da Cabeceira, on flowers of Oenanthe crocata L. by day, 25-V-2019, Corley.

Micropterix ibericella Caradja, 1920

R: Valverde, by day, 24-V-2019, S. Ferreira.

NEPTICULIDAE

Stigmella basiguttella (Heinemann, 1862)

DL: Couce, leafmine on Quercus róbur L., 18-XI-2019, Nunes.

Stigmella atricapitella (Haworth, 1828)

TM: Lordelo, 11-VII-2019, Gonzalez, det. Corley.

Trifurcula orientella Klimesch, 1953

BL: Outeiro dos Gamanhos, 28-VI-2019, Rosete, det. Corley.

Bohemannia pulverosella (Stainton, 1849)

Second Portuguese record. TM: Dine, empty leafmine on Malus sp., 20-IX-2019, Corley and Nunes.

Ectoedemia suberis (Stainton, 1869)

DL: Valongo, leafmine on Quercus suber L., 1-III-2019, Nunes, det. Corley from reared adult.

TISCHERIIDAE

Coptotriche marginea (Haworth, 1828)

BAL: Moinhos de Paneiro, 16-IV-2019, Nunes, Silva and Jesus.

ERIOCOTTIDAE

Eriocottis hispanica Zagulajev, 1988

BA: Faia Brava, 10-V-2019, Nunes, Silva and Jesus.

PSYCHIDAE

Placodoma calpella Sobczyk, 2013

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira, det. Arnscheid.

Ptilocephala albida (Esper, 1786)

BL: Buracas de Casmilo, 1-V-2019, by day, Rosete.

Delete Phalacropterix graminifera (Fourcroy, 1785) from Portuguese list. A specimen in SMNK collected by T. Monteiro and determined as the synonym constancella Bruand, 1853 by Sieder is P. fritschi Hättenschwiler, 2003 (W. Arnscheid, pers. comm.). Other Monteiro specimens of P. graminiferacan be assumed to belong to P. fritschi, see below.

Phalacropterix fritschi Hättenschwiler, 2003

= constancella sensu Monteiro, 1956

DL: Singeverga, VI-1948, Monteiro (SMNK). Monteiro’s other records of constancella from Mosteiro de Singeverga, Citânia de Sanfins, Santa do Amparo, Azurara and Pedras Rubras (MONTEIRO, 1956) are all from Douro Litoral. As Monteiro would have named these by comparison with the Singeverga specimen named by Sieder, there can be little doubt that all belong to P. fritschi. This is a considerable extension of range for the Portuguese endemic P. fritschi, previously known only from Algarve.

TINEIDAE

Morophaga morella (Duponchel, 1838)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Nemapogon agenjoi Petersen, 1959

R: Colos, 13-VI-2019, Terry.

Triaxomasia caprimulgella (Stainton, 1851)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Neurothaumasia ankerella (Mann, 1867)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Reisserita zernyi Petersen, 1957

R: Entrevinhas, Sardoal, 25-VII-2019, Rosete and Lameirinhas.

Monopis obviella (Denis & Schiffermüller, 1775)

TM: Guadramil, 20-IX-2019, Corley, Nunes and S. Ferreira.

Crassicornella agenjoi Petersen, 1957

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Ateliotum petrinella (Herrich-Schäffer, 1854)

ALG: Corte de Pero Jaques, 24-VIII-2019, Rosete, det. Corley.

BUCCULATRICIDAE

Bucculatrix albedinella (Zeller, 1839)

Second Portuguese record. TM: Guadramil, leafmines on Ulmus minor Mill., 20-IX-2019, Corley and Nunes.

Bucculatrix diffusella Menhofer, 1943

BL: Praia do Cabedelo, 16-VII-2019, Rosete, det. Corley.

GRACILLARIIDAE

Parectopa ononidis (Zeller, 1839)

BAL: Canto Mosqueiro, 28-V-2019, Corley and S. Ferreira.

Micrurapteryx kollariella (Zeller, 1839)

BA: Covão da Ponte, 16-VIII-2019, Rosete.

Euspilapteryx auroguttella Stephens, 1835

R: Colos, 10-VI-2019, Terry.

Spulerina simplonia (Fischer von Röslerstamm, 1840)

R: Colos, 10-VI-2019, Terry.

Phyllonorycter mespilella (Hübner, 1805)

M: Curras Grandes, leafmine on Pyrus cordata Desv., 3-VI-2019, Corley.

Phyllonorycter floridae Laštůvka & Laštůvka, 2012

TM: Vimioso, 8-VIII-2019, Gonzalez, det. Corley.

* Phyllonorycter comparella (Duponchel, 1843)

TM: R. Angueira below Castelo de Algoso, leafmines on Populus canescens (Aiton) Sm., 24-IX- 2019, Corley. Widespread in central Europe, also present in a few countries further north and south and in some Mediterranean Islands. [Also recorded from Populus alba L. and P. nigra L.].

YPONOMEUTIDAE

Zelleria oleastrella (Millière, 1864)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira; R: Colos, 8-VI-2019, Terry.

Swammerdamia pyrella (Villers, 1789)

M: Curras Grandes, larva on Pyrus cordata Desv., 3-VI-2019, Corley.

PRAYDIDAE

Prays oleae (Bernard, 1788)

R: Outeiro, Alcobertas, 23-V-2019, Corley.

DOUGLASIIDAE

Tinagma ocnerostomella (Stainton, 1850)

E: Rua do Furo, 23-V-2019, Corley and S. Ferreira.

AUTOSTICHIDAE

Arragonia punctivittella (Zerny, 1927)

TM: Guadramil, 17-VIII-2019, Nunes, Silva and Jesus.

Oegoconia novimundi (Busck, 1915)

TM: São Martinho de Angueira, 23-IX-2019, Corley and S. Ferreira.

** Apatema apolausticum Gozmány, 1996

ALG: Serra de Monte Figo, 19-V-2002, Corley. South-east Europe from Italy and Slovakia to Romania. [Host-plant unknown but probably feeds on plant detritus].

Symmocoides oxybiella (Millière, 1872)

BL: Ansião, 16-VII-2019, Rosete.

Stibaromacha ratella (Herrich-Schäffer, 1854)

R: Colos, 10-VI-2019, Terry.

LECITHOCERIDAE

Homaloxestis briantiella (Turati, 1879)

BL: Ansião, 16-VII-2019, Rosete, det. Corley.

Eurodachtha pallicornella (Staudinger, 1859)

R: Colos, 13-VI-2019, Terry.

OECOPHORIDAE

Kasyniana griseosericeella (Ragonot, 1879)

BAL: Vale de Gomes, 27-V-2019, Corley and S. Ferreira.

Alabonia herculeella Walsingham, 1903

E: Quinta do Pisão, 12-V-2019, Tagis, det. Corley.

Pleurota andalusica Back, 1973

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira; R: Colos, 10-VI-2019, Terry; BL: Ansião, 16-VII-2019, Rosete, det. Corley.

Pleurota planella (Staudinger, 1859)

BL: Abiúl, 31-V-2019, Corley and Rosete.

DEPRESSARIIDAE

Agonopterix nodiflorella (Millière, 1866)

E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira.

Agonopterix umbellana (Fabricius, 1794)

BL: Buracas de Casmilo, 2-VIII-2019, Rosete.

Agonopterix heracliana (Linnaeus, 1758)

E: Vale da Cabeceira, larvae on flowers of Oenanthe crocata L., 25-V-2019, Corley.

Agonopterix thapsiella (Zeller, 1847)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Agonopterix alstromeriana (Clerck, 1759)

DL: Valongo, 24-I-2019, Nunes.

Agonopterix subpropinquella (Stainton, 1849)

R: Valverde, larvae on Leuzea conifera (L.) DC, 24-V-2019, Corley.

Exaeretia lutosella (Herrich-Schäffer, 1854)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira; R: Colos, 8-VI-2019, Terry.

Depressaria daucella (Denis & Schiffermüller, 1775)

E: Vale da Cabeceira, larvae on flowers of Oenanthe crocata L., 25-V-2019, Corley.

COSMOPTERIGIDAE

Eteobalea intermediella (Riedl, 1966)

R: Colos, 8-VI-2019, Terry.

Vulcaniella fiordalisa (Petry, 1904)

BAL: Vale de Gomes, 27-V-2019, Corley and S. Ferreira.

GELECHIIDAE

Aproaerema sangiella (Stainton, 1863)

E: Azelha, 13-VI-2019, Terry; BAL: Canto Mosqueiro, 28-V-2019, Corley and S. Ferreira.

Mesophleps oxycedrella (Millière, 1871)

E: Azelha, 13-VI-2019, Terry.

Nothris congressariella (Bruand, 1858)

BL: Abiúl, 31-V-2019, Corley and Rosete.

Neofaculta ericetella (Geyer, 1832)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Dichomeris acuminatus (Staudinger, 1876)

BAL: Canto Mosqueiro, 28-V-2019, Corley and S. Ferreira.

Dichomeris limbipunctella (Staudinger, 1859)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Catatinagma agenjoi (Gozmány, 1954)

Coloptilia conchylidella (Hofmann, 1898) was recorded in Algarve in CARVALHO & CORLEY (1995), but subsequently excluded from the Portuguese fauna in CORLEY (2015) without a correct identification for the specimen being provided. Recent re-examination shows that it belongs to Catatinagma agenjoi.

Bryotropha terrella (Denis & Schiffermüller, 1775)

E: Rua do Furo, 23-V-2019, Corley and S. Ferreira.

Aristotelia subdecurtella (Stainton, 1859)

BAL: Lagoa de Santo André, 26-V-2019, Corley.

Aristotelia brizella (Treitschke, 1833)

BAL: Lagoa de Santo André, 26-V-2019, Corley.

Isophrictis anthemidella (Wocke, 1871)

E: Vale da Cabeceira, 25-V-2019, Corley; BA: Fráguas, 23-VII-2019, Tagis, det. Corley.

Metzneria aestivella (Zeller, 1839)

R: Colos, 8-VI-2019, Terry.

Metzneria littorella (Douglas, 1850)

BL: Peixeiro, 4-V-2019, Rosete, det. Corley.

Metzneria tristella Rebel, 1901

BL: Abiúl, 22-VI-2019, Rosete, det. Corley.

Ptocheuusa paupella (Zeller, 1847)

R: Colos, 8-VI-2019, Terry.

Oxypteryx immaculatella (Douglas, 1850)

R: Colos, 10-VI-2019, Terry.

Oxypteryx helotella (Staudinger, 1859)

BAL: Lagoa de Santo André, 26-V-2019, Corley.

* Gladiovalva rumicivorella (Millière, 1881)

M: Serra de Arga, 3-VIII-2019, Silva and Jesus (fig. 1). Spain to Germany, Switzerland and Italy. [Rumex scutatus L.].

Mirificarma denotata Pitkin, 1984

E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira.

Psoricoptera gibbosella (Zeller, 1839)

TM: Carrazedo, 18-IX-2019, Corley, Nunes & S. Ferreira.

Scrobipalpa acuminatella (Sircom, 1850)

BL: Madriz, 5-VII-2019, Rosete, det. Corley.

Scrobipalpa vasconiella (Rössler, 1877)

R: Colos, 9-VI-2019, Terry.

Scrobipalpa artemisiella (Treitschke, 1833)

TM: Guadramil, 17-VIII-2019, Nunes, Silva and Jesus, det. Corley.

Scrobipalpa salinella (Zeller, 1847)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Tuta absoluta (Meyrick, 1917)

R: Valverde, 24-V-2019, Corley and S. Ferreira; E: Vale da Cabeceira, 25-V-2019, Corley.

Klimeschiopsis terroris (Hartig, 1938)

TM: Estevais, 24-IX-2019, Corley and S. Ferreira.

Teleiodes albidorsella Huemer & Karsholt, 1999

M: Vale de Branco, 27-VIII-2019, Tagis, det. Corley.

Teleopsis diffinis (Haworth, 1828)

R: Colos, 10-VI-2019, Terry; E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira.

Pseudotelphusa scalella (Scopoli, 1763)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Pseudotelphusa occidentella Huemer & Karsholt, 1999

BAL: Moinhos de Paneiro, 16-IV-2019, Nunes, Silva and Jesus.

Recurvaria leucatella (Clerck, 1759)

BL: Buracas de Casmilo, 2-VIII-2019, Rosete.

ELACHISTIDAE

Elachista hispanica Traugott-Olsen, 1992

R: Colos, 10-VI-2019, Terry.

Elachista obliquella Stainton, 1854

E: Vale da Cabeceira, 25-V-2019, Corley.

Haplochrois ochraceella (Rebel, 1903)

E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira.

COLEOPHORIDAE

Coleophora conyzae Zeller, 1868

R: Colos, 10-VI-2019, Terry.

Coleophora acrisella Millière, 1872

Recorded feeding on Coronilla glauca (CORLEY, 2015), but these records are misidentifications of Coleophora sisteronica Toll, 1961. Other records are correct.

Coleophora sisteronica Toll, 1961

ALG: Silves, cases on Coronilla glauca L., 9-XI-2000, 18-III-2011, Corley; third record, BL: Abiúl, cases on Coronilla repanda (Poir.) Guss., 31-V-2019, Corley and Rosete.

Coleophora deauratella Lienig & Zeller, 1846

R: Colos, 13-VI-2019, Terry.

Coleophora amethystinella Ragonot, 1885

BA: Vilar do Amargo, 11-V-2019, Nunes, det. Corley.

** Coleophora currucipennella Zeller, 1839

TM: Pitões das Júnias, 24-VII-2019, Nunes and Jesus, det. Corley (fig. 2). Middle latitudes of Europe, reaching north to southern parts of Scandinavia and south to a few Mediterranean countries, including Greece. [Quercus sp., Salix sp., Carpinus sp.].

Coleophora helichrysiella Krone, 1909

BAL: Monte Velho, 26-V-2019, Corley and S. Ferreira.

Coleophora albicosta (Haworth, 1828)

E: Rua do Furo, 23-V-2019, Corley and S. Ferreira.

Coleophora pennella (Denis & Schiffermüller, 1775)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Coleophora alticolella Zeller, 1849

BA: Covão da Ametade, 9-VII-2019, Rosete, det. Corley.

Coleophora maritimella Newman, 1863

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

* Coleophora ribasella Baldizzone, 1982

BL: Praia do Cabedelo, 31-VIII-2017, reared from case on Artemisia campestris maritima (DC) Arcag, Rosete, det. S. Ferreira and Corley. [Spain, France].

SCYTHRIDIDAE

Scythris subseliniella (Heinemann, 1876)

BA: Lagoa Comprida, 9-VII-2019, Rosete, det. Corley.

Scythris potentillella (Zeller, 1847)

BA: Vilar de Amargo, 11-V-2019, Nunes, Silva and Jesus, det. Corley; TM: Turning to Lama Grande, 14-VIII-2019, Tagis, det. Corley.

Scythris cistorum (Millière, 1876)

BAL: Foros da Quinta, 26-V-2019, Corley and S. Ferreira.

Scythris scopolella (Linnaeus, 1767)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Scythris lafauryi Passerin d’Entrèves, 1986

TM: Santo André, 16-IX-2019, Corley, Nunes and S. Ferreira.

Episcythris triangulella (Ragonot, 1874)

BL: Relvas, 26-VII-2019, Rosete, det. Corley.

BLASTOBASIDAE

Blastobasis phycidella (Zeller, 1839)

DL: Vila Nova de Gaia, 25-VI-2010, Corley.

* Hypatopa segnella (Zeller, 1873)

TM: Alto de Ciradelhe, 10-VIII-2019, Tagis, det. Corley. Distribution very scattered from South Russia and Finland to France and Spain, but absent from many countries. [Larva on plant detritus].

MOMPHIDAE

Mompha miscella (Denis & Schiffermüller, 1775)

BAL: Foros da Quinta, 26-V-2019, Corley and S. Ferreira.

Mompha propinquella (Stanton, 1851)

BA: Covão da Ponte, 16-VIII-2019, Rosete.

PTEROLONCHIDAE

Pterolonche lutescentella Chrétien, 1922

BL: Buracas de Casmilo, 2-VIII-2019, Rosete, det. Corley.

LYPUSIDAE

Agnoea amparoella (Vives, 1986)

BAL: Vale de Gomes, 27-V-2019, Corley and S. Ferreira; E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira.

ALUCITIDAE

Alucita hexadactyla Linnaeus, 1758

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Alucita huebneri Wallengren, 1859

BL: Ansião, 20-V-2017, Rosete, det. Corley.

PTEROPHORIDAE

Agdistis meridionalis (Zeller, 1847)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Stenoptilia pelidnodactyla (Stein, 1857)

BB: Cântaro Raso, 24-VI-2010, Corley.

Stenoptilia zophodactylus (Duponchel, 1840)

R: Colos, 8-VI-2019, Terry.

Merrifieldia tridactyla (Linnaeus, 1758)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Merrifieldia baliodactylus (Zeller, 1841)

BL: Chão do Ulmeiro, 27-VI-2019, Rosete, det. Corley.

Merrifieldia malacodactylus (Zeller, 1847)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

SCHRECKENSTEINIIDAE

* Schreckensteinia festaliella (Hübner, 1796)

E: Vale da Cabeceira, 25-V-2019, Corley. Almost all Europe, but absent from Mediterranean islands and some countries in the south-east. [Rubus fruticosus L., R. idaeus L.].

EPERMENIIDAE

Epermenia aequidentellus (E. Hofmann, 1867)

E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira; R: Colos, 8-VI-2019, Terry.

Epermenia ochreofasciella (Millière, 1854)

Third and fourth records. TM: Balsamão, 22-IX-2019, Corley and S. Ferreira; São Martinho da Angueira, 23-IX-2019, Corley and S. Ferreira.

CHOREUTIDAE

Tebenna micalis (Mann, 1857)

R: Colos, 9-VI-2019, Terry.

Choreutis nemorana (Hübner, 1799)

BA: Penhas Douradas, 16-VIII-2019, Rosete.

TORTRICIDAE

Isotrias penedana Trematerra, 2013

TM: Marão, netted by day, 15-VI-2019, Nunes, Silva and Jesus (fig. 3). This male differs from those found in the Castro Laboreiro area (TREMATERRA, 2013) in having indications of the yellow transverse markings that are found in the females (CORLEY & FERREIRA, 2017), although still with a reticulate pattern over the whole forewing. Male genitalia show no differences from those figured by TREMATERRA (2013).

Paramesia alhamana (Schmidt, 1933)

E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira; BL: Abiúl, 31-V-2019, Corley and Rosete.

Avaria joannisiana (Ragonot, 1888)

ALG: Sargaçal, 13-XI-2019, Valadares, det. Corley.

Syndemis musculana (Hübner, 1799)

DL: Vale Quente, Serra do Marão, 15-VI-2019, Nunes, Silva and Jesus.

Lozotaenia cupidinana (Staudinger, 1859)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Clepsis coriacana (Rebel, 1894)

BAL: Canto Mosqueiro, larva on Lotus creticus, 28-V-2019, Corley and S. Ferreira.

Clepsis peritana (Clemens, 1860)

BAL: Canto Mosqueiro, 28-V-2019, Corley and S. Ferreira; BL: Figueiró do Campo, 31-VIII- 2019, I. Ferreira.

Tortricodes alternella (Denis & Schiffermüller, 1775)

BA: Picão, 4-I-2019, Nunes, Silva and Jesus.

Cnephasia stephensiana (Doubleday, 1849)

E: Rua do Furo, 23-V-2019, Corley and S. Ferreira.

Cnephasia longana (Haworth, 1811)

E: Berlenga,14-V-2019, Tagis, det. Corley; BL: Buracas de Casmilo, 1-V-2019, Rosete, det. Corley; Curras Grandes, 3-VI-2019, Corley.

Cnephasia ecullyana Réal, 1951

BAL: Vale de Gomes, 27-V-2019, Corley and S. Ferreira.

Tortrix viridana Linnaeus, 1758

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Aleimma loeflingiana (Linnaeus, 1758)

R: Outeiro, Alcobertas, 23-V-2019, Corley.

Acleris schalleriana (Linnaeus, 1761)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira; BL: Abiúl, 31-V-2019, Corley and Rosete.

Cochylimorpha elongana (Fischer von Röslerstamm, 1839)

BAL: Lagoa de Santo André, 26-V-2019, Corley and S. Ferreira.

Agapeta angelana (Kennel, 1919)

Second Portuguese record. BL: São Romão, Vagos, 10-VI-2019, Tagis, det. Corley.

Eugnosta lathoniana (Hübner, 1800)

TM: Guadramil, 17-VIII-2019, Nunes, Silva and Jesus (fig. 4). Only one previous record MENDES (1910).

Aethes beatricella (Walsingham, 1898)

BL: Ansião, 16-VII-2019, Rosete, det. Corley.

Aethes bilbaensis (Rössler, 1877)

R: Colos, 9-VI-2019, Terry.

* Aethes mauritanica (Walsingham, 1898)

BAL: Vila Nova de Milfontes, 27-V-2019, Corley and S. Ferreira. South Europe from Spain and Corsica to Greece and Bulgaria. [Elaeoselinum meoides (Desf.) W. D. J. Koch ex DC].

Diceratura infantana (Kennel, 1899)

BAL: Monte Velho, Lagoa de Santo André, 26-V-2019, Corley and S. Ferreira.

** Neocochylis millierana (Peyerimhoff, 1877)

TM: Cigadonhe, Gestosa, 16-VIII-2019, Nunes, Silva and Jesus, det. Corley (fig. 5). NEL & VARENNE (2017) show that millierana is not synonymous with salebrana, which has one record from Algarve (CORLEY, 2005), but with Cochylis sannitica Trematerra, 1995, known from France and Italy. Host-plant unknown.

Neocochylis molliculana (Zeller, 1847)

BAL: Lagoa de Santo André, 26-V-2019, Corley.

Brevicornutia pallidana (Zeller, 1847)

BAL: Lagoa de Santo André, 26-V-2019, Corley.

Piniphila bifasciana (Haworth, 1811)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Lobesia bicinctana (Duponchel, 1844)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Lobesia artemisiana (Zeller, 1847)

E: Berlenga,14-V-2019, Tagis, det. Corley.

Endothenia gentianaeana (Hübner, 1799)

BA: Nodar, Serra de Montemuro, 22-VII-2019, Tagis, det. Corley.

Endothenia marginana (Haworth, 1811)

BAL: Lagoa de Santo André, 26-V-2019, Corley.

Ancylis obtusana (Haworth, 1811)

DL: Vale Quente, Serra do Marão, 15-VI-2019, Nunes, Silva and Jesus.

Thiodia trochilana (Frölich, 1828)

E: Berlenga,14-V-2019, Tagis, det. Corley.

Acroclita subsequana (Herrich-Schäffer, 1851)

R: Outeiro, Alcobertas, 23-V-2019, Corley.

Epinotia bilunana (Haworth, 1811)

Third Portuguese locality. TM: Quintã, 20-V-2016, Fernandes and Gonzalez, det. Corley.

Zeiraphera isertana (Fabricius, 1794)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Gypsonoma imparana Müller-Rutz, 1914

M: Serra de Arga: Parque Eólico, larva on Salix atrocinerea, 3-VI-2019, Corley.

Notocelia incarnatana (Hübner, 1800)

BAL: Foros da Quinta, 26-V-2019, Corley and S. Ferreira.

Notocelia trimaculana (Haworth, 1811)

E: Rua do Furo, 23-V-2019, Corley and S. Ferreira.

* Pseudococcyx friedmariana Larsen, 2020

AAL: Minhota, 5-VI-1996, Corley; Marco, E. of Esperança, 12-V-1999, Corley; BL: Ceira, 25-V- 2006, Pires, det. Corley; Poço, 7-VI-2016, Rosete, det. Corley (fig. 6); BAL: Praia do Malhão, 12-V- 2017, Marabuto, Grundy, Nunes, Silva and Jesus, det. Corley; Lagoa de Santo André, 26-V-2019 Corley and S. Ferreira. The species was recently described from two localities in Spain, based on two male specimens (LARSEN, 2020). In Portugal only females have been found. These have been barcoded but barcodes are not available for Spanish material. The conspecificity of Spanish and Portuguese material is therefore not proven, but we consider it probable. Host-plant unknown.

Delete Retinia resinella (Linnaeus, 1758) from Portuguese list. The record and photo from BAL: Praia do Malhão in CORLEY et al. (2018) clearly belong to Pseudococcyx friedmariana (above). R. resinella has only two other Portuguese records: BL: Mata de Leiria, A. F. de Seabra (SEABRA, 1939) and ALG: São Romão, V-1982, J. Passos de Carvalho (MONTEIRO & CARVALHO, 1984). Without supporting evidence and in view of the possibility of confusion of this species with P. friedmariana, these two isolated records must be considered doubtful. The species should therefore be removed from the Portuguese list.

Clavigesta sylvestrana (Curtis, 1850)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Rhyacionia buoliana (Denis & Schiffermüller, 1775)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Cydia gilviciliana (Staudinger, 1859)

BAL: Canto Mosqueiro, 28-V-2019, Corley and S. Ferreira.

BRACHODIDAE

Brachodes gaditana (Rambur, 1866)

BL: Poço, 18-VII-2019, Rosete.

COSSIDAE

Dyspessa ulula (Borkhausen, 1790)

BAL: Noudar, Barrancos, 10-IV-2019, Tagis; R: Colos, 10-VI-2019, Terry.

SESIIDAE

Bembecia ichneumoniformis (Denis & Schiffermüller, 1775)

R: Golegã, 12-VI-2019, Terry.

Bembecia uroceriformis (Treitschke, 1834)

BL: Outeiro dos Gamanhos, 29-VI-2019, Rosete, det. Corley; TM: Guadramil, 17-VIII-2019, Nunes, Silva and Jesus.

Pyropteron meriaeformis (Boisduval, 1840)

E: Marateca, 5-VI-2019, Tagis, det. Corley.

PYRALIDAE

Pyralis lienigialis (Zeller, 1843)

BAL: Vale de Gomes, 27-V-2019, Corley and S. Ferreira; E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira.

Bostra obsoletalis (Mann, 1864)

R: Colos, 8-VI-2019, Terry.

Stemmatophora combustalis (Fischer von Röslerstamm, 1842)

R: Colos, 13-VI-2019, Terry.

Stemmatophora rungsi (Leraut, 2000)

DL: Regoufe, 21-VII-2019, Tagis, det. Corley.

Synaphe moldavica (Esper, 1794)

BL: Chão do Ulmeiro, 27-VI-2019, Rosete.

Cryptoblabes gnidiella (Millière, 1867)

TM: Alto de Ciradelhe, 10-VIII-2019, Tagis, det. Corley.

Bradyrrhoa cantenerella (Duponchel, 1837)

BL: Vale da Couda, 4-VII-2019, Rosete.

Pempelia palumbella (Denis & Schiffermüller, 1775)

R: Colos, 10-VI-2019, Terry.

Asalebria florella (Mann, 1862)

E: Rua do Furo, 23-V-2019, Corley and S. Ferreira; R: Colos, 9-VI-2019, Terry.

Dioryctria sylvestrella (Ratzeburg, 1840)

TM: Lordelo, 2-IX-2016, Fernandes and Gonzalez, det. Corley.

Dioryctria mendacella (Staudinger, 1859)

R: Colos, 12-VI-2019, Terry.

Epischnia prodromella (Hübner, 1799)

M: Mosteiro de São João de Arga, 25-IX-2019, Corley and S. Ferreira.

Elegia fallax (Staudinger, 1881)

= fallaximima Nel & Mazel, 2011; fallax sensu Slamka, 2019

BAL: Vale de Gomes, 27-V-2019, Corley and S. Ferreira; E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira.

Elegia similella (Zincken, 1818)

TM: Pitões das Júnias, 24-VII-2019, Nunes and Jesus.

Pima boisduvaliella (Guenée, 1845)

BAL: Praia dos Aivados, 10-III-2019, Tagis, det. Corley.

Melathrix coenulentella (Zeller, 1846)

R: Colos, 13-VI-2019, Terry.

Delplanqueia dilutella (Denis & Schiffermüller, 1775)

DL: Regoufe, 21-VII-2019, Tagis, det. Corley.

Delplanqueia inscriptella (Duponchel, 1836)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Pempeliella ardosiella (Ragonot, 1887)

BAL: Foros da Quinta, 26-V-2019, Corley and S. Ferreira.

Psorosa mediterranella Amsel, 1953

BL: Quinta do Canal, 14-IX-2019, Corley, Rosete, Nunes and S. Ferreira.

Acrobasis obliqua (Zeller, 1847)

R: Outeiro, Alcobertas, 23-V-2019, Corley.

Acrobasis consociella (Hübner, 1813)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Acrobasis suavella (Zincken, 1818)

R: Colos, 10-VI-2019, Terry; BL: Ansião, 16-VII-2019, Rosete.

Acrobasis marmorea (Haworth, 1811)

R: Colos, 9-VI-2019, Terry; BA: Vilar de Amargo, 11-V-2019, Nunes, Silva and Jesus.

Myelois circumvoluta (Fourcroy, 1785)

R: Colos, 13-VI-2019, Terry.

Apomyelois ceratoniae (Zeller, 1839)

R: Ave Casta, 21-V-2019, Tagis, det. Corley.

Eccopisa effractella Zeller, 1848

E: Vale da Cabeceira, 25-V-2019, Corley.

Euzophera fuliginosella (Heinemann, 1865)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Nyctegretis ruminella La Harpe, 1860

BAL: Vale de Gomes, 27-V-2019, Corley and S. Ferreira.

Ancylosis cinnamomella (Duponchel, 1836)

R: Outeiro, Alcobertas, 23-V-2019, Corley.

Ancylosis sareptalla (Herrich-Schäffer, 1861)

BAL: Vila Nova de Milfontes, 28-V-2019, Corley and S. Ferreira.

Ancylosis arenosella (Staudinger, 1859)

BAL: Praia dos Aivados, 10-III-2019, Tagis, det. Corley.

Phycitodes inquinatella (Ragonot, 1887)

R: Colos, 10-VI-2019, Terry.

Ephestia welseriella (Zeller, 1848)

R: Colos, 12-VI-2019, Terry.

CRAMBIDAE

Pyrausta sanguinalis (Linnaeus, 1767)

R: Outeiro, Alcobertas, 23-V-2019, Corley.

Pyrausta neglectalis Caradja, 1916

R: Colos, 9-VI-2019, Terry.

Pyrausta despicata (Scopoli, 1753)

R: Colos, 12-VI-2019, Terry.

Anania terrealis (Treitschke, 1829)

TM: Serra do Larouco, 15-IX-2019, Corley, Nunes and S. Ferreira.

Anania verbascalis (Denis & Schiffermüller, 1775)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Udea ferrugalis (Hübner, 1796)

R: Colos, 8-VI-2019, Terry.

Udea numeralis (Hübner, 1796)

R: Colos, 8-VI-2019, Terry.

Mecyna asinalis (Hübner, 1819)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Agrotera nemoralis (Scopoli, 1763)

ALG: Alfambras, 7-VIII-2019, Valadares.

Palpita vitrealis (Rossi, 1794)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Dolicharthria aetnealis (Duponchel, 1833)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Cynaeda dentalis (Denis & Schiffermüller, 1775)

R: Colos, 13-VI-2019, Terry; E: Berlenga,14-V-2019, Tagis, det. Corley.

Evergestis politalis (Denis & Schiffermüller, 1775)

R: Outeiro, Alcobertas, 23-V-2019, Corley.

Scoparia pyralella (Denis & Schiffermüller, 1775)

DL: Vale Quente, Serra do Marão, 15-VI-2019, Nunes, Silva and Jesus.

Scoparia staudingeralis (Mabille, 1869)

BAL: Safara, 12-IV-2019, Tagis, det. Corley.

Eudonia angustea (Curtis, 1827)

R: Olhos de Água de Alviela, 18-V-2019, Tagis, det. Corley.

Eudonia lineola (Curtis, 1827)

R: Outeiro, Alcobertas, 23-V-2019, Corley.

Eudonia delunella (Stainton, 1849)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Heliothela wulfeniana (Scopoli, 1763)

BB: Cortes de Meio, 22-VI-2019, Tagis, det. E. Marabuto.

Chrysoteuchia culmella (Linnaeus, 1758)

BAL: Lagoa de Santo André, 26-V-2019, Corley.

Agriphila deliella (Hübner, 1813)

M: Mosteiro de São João de Arga, 25-IX-2019, Corley and S. Ferreira.

Agriphila tersellus (Lederer, 1855)

BL: Quinta do Canal, 14-IX-2019, Corley, Rosete, Nunes and S. Ferreira.

* Catoptria lythargyrella (Hübner, 1796)

TM: Montesinho, 19-IX-2019, Corley, Nunes and S. Ferreira. Middle latitudes of Europe, absent from several countries in north and south and some Mediterranean islands. [Grasses].

Mesocrambus tamsi Bleszynski, 1960

BA: Escalhão, 10-VII-2019, Tagis, det. Corley.

Hyperlais siccalis (Guenée, 1854)

R: Outeiro, Alcobertas, 23-V-2019, Corley; E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira.

Elophila nymphaeata (Linnaeus, 1758)

BA: Vilar de Amargo, 11-V-2019, Nunes, Silva and Jesus.

LASIOCAMPIDAE

Macrothylacia digramma Meade-Waldo, 1905

R: Valverde, 24-V-2019, Corley and S. Ferreira.

SATURNIIDAE

Saturnia pavonia (Linnaeus, 1758)

BAL: Praia dos Aivados, 10-III-2019, Tagis.

SPHINGIDAE

Marumba quercus (Denis & Schiffermüller, 1775)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Mimas tiliae (Linnaeus, 1758)

E: Sintra (CARNEIRO MENDES, 1951). Accidentally omitted by CORLEY (2015).

Sphinx ligustri Linnaeus, 1758

R: Sentieiras, 20-V-2019, Érica Brunheta, comm. A. Lameirinhas.

Hyles euphorbiae (Linnaeus, 1758)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Deilephila porcellus (Linnaeus, 1758)

BB: Parque Campismo de Pião, 10-VI-2019, Nunes, Silva and Jesus.

GEOMETRIDAE

Idaea sardoniata (Homberg, 1912)

R: Colos, 8-VI-2019, Terry; E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira.

Idaea consanguiberica Rezbanyai-Reser & Expósito, 1992

BA: Castelo Bom, 12-VIII-2019, Skou.

Idaea mustelata (Gumppenberg, 1892)

BAL: Vila Nova de Milfontes, 28-V-2019, Corley and S. Ferreira.

Idaea efflorata Zeller, 1849

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Idaea incisaria (Staudinger, 1892)

BA: Vilar de Amargo, 11-V-2019, Nunes, Silva and Jesus.

Idaea robiginata (Staudinger, 1863)

DL: Vale Quente, Serra do Marão, 15-VI-2019, Nunes, Silva and Jesus.

Idaea bigladiata Herbulot, 1975

R: Outeiro, Alcobertas, 23-V-2019, Corley; BA: Faia Brava Reserve, 10-V-2019, Nunes, Silva and Jesus.

Idaea carvalhoi Herbulot, 1979

R: Colos, 8-VI-2019, Terry.

Idaea dimidiata (Hufnagel, 1767)

E: Rua do Furo, 23-V-2019, Corley and S. Ferreira.

Idaea ostrinaria (Hübner, 1813)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Idaea dromikos Hausmann, 2004

BA: Covão da Ponte, 16-VIII-2019, Rosete.

Idaea straminata (Borkhausen, 1794)

R: Colos, 10-VI-2019, Terry.

Anthometra plumularia Boisduval, 1840

ALG: Alfambras, 2-VII-2016, 28-VI-2018, Valadares; R: Colos, 9-VI-2019, Terry.

Catarhoe rubidata (Denis & Schiffermüller, 1775)

TM: Pitões das Júnias, 15-VII-2019, Nunes and Pires.

Larentia clavaria (Haworth, 1809)

BL: Moura da Serra, 26-X-2019, Rosete.

Euphyia unangulata (Haworth, 1809)

Second Portuguese record. M: Mosteiro de São João de Arga, 3-VI-2019, Corley and S. Ferreira.

Thera obeliscata (Hübner, 1787)

DL: Vale Quente, Serra do Marão, 15-VI-2019, Nunes, Silva and Jesus.

Chloroclysta miata (Linnaeus, 1758)

TM: Montesinho, 19-IX-2019, Corley, Nunes and S. Ferreira.

Asthena albulata (Hufnagel, 1767)

BL: Moura da Serra, 15-V-2016, Rosete. The record for BL given in CORLEY (2015) belongs to DL.

Chloroclystis v-ata (Haworth, 1809)

E: Rua do Furo, 23-V-2019, Corley and S. Ferreira.

Eupithecia cocciferata Millière, 1864

BAL: Moinhos de Paneiro, 16-IV-2019, Nunes, Silva and Jesus.

Eupithecia abbreviata Stephens, 1831

E: Quinta do Pisão, 12-V-2019, Tagis, det. Corley.

Eupithecia dodoneata Guenée, 1858

R: Colos, 9-VI-2019, Terry.

Eupithecia massiliata Millière, 1863

BAL: Moinhos de Paneiro, 16-IV-2019, Nunes, Silva and Jesus.

Eupithecia scopariata (Rambur, 1833)

R: Colos, 9-VI-2019, Terry.

Eupithecia breviculata (Donzel, 1837)

BAL: Foros da Quinta, 26-V-2019, Corley and S. Ferreira; R: Colos, 8-VI-2019, Terry.

* Eupithecia assimilata Doubleday, 1856

TM: Penelas, Ermida, larva on Humulus lupulus L., 8-X-2016, Nunes, conf. Townsend (fig. 7). Almost all Europe, absent from some countries in south-east. [Larva also on Ribes sp.].

Aplocera efformata (Guenée, 1858)

BAL: Vila Nova de Milfontes, 28-V-2019, Corley and S. Ferreira.

Chesias rufata (Fabricius, 1775)

BAL: Moinhos de Paneiro, 16-IV-2019, Nunes, Silva and Jesus.

Acasis viretata (Hübner, 1799)

BAL: Santa Bárbara, 11-III-2019, Tagis.

Lomaspilis marginata (Linnaeus, 1758)

BL: Jafafe de Cima, 12-VI-2019, Tagis.

Acanthovalva inconspicuaria (Hübner, 1819)

E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira.

Neognopharmia stevenaria (Boisduval, 1840)

BAL: Noudar, Barrancos, 10-IV-2019, Tagis.

Petrophora chlorosata (Scopoli, 1763)

BB: Parque Campismo de Pião, 10-VI-2019, Nunes, Silva and Jesus.

Selenia lunularia (Hübner, 1788)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira.

Tephronia espaniola Schawerda, 1931

Second Portuguese record. TM: Cigadonhe, Gestosa, 16-VIII-2019, Nunes, Silva and Jesus (fig. 8).

Hylaea fasciaria (Linnaeus, 1758)

DL: Vale Quente, Serra do Marão, 15-VI-2019, Nunes, Silva and Jesus.

Charissa crenulata avilarius (Reisser, 1936)

BB: Parque Campismo de Pião, 10-VI-2019, Nunes, Silva and Jesus.

Thalera fimbrialis (Scopoli, 1763)

R: Colos, 8-VI-2019, Terry.

NOTODONTIDAE

Stauropus fagi (Linnaeus, 1758)

BAL: Santa Bárbara, 11-III-2019, Tagis.

Notodonta dromedarius (Linnaeus, 1767)

DL: Vale Quente, Serra do Marão, 15-VI-2019, Nunes, Silva and Jesus.

Ptilodon capucina (Linnaeus, 1758)

Third Portuguese record. M: Mosteiro de São João de Arga, 3-VI-2019, Corley and S. Ferreira.

EREBIDAE

Orgyia aurolimbata Guenée, 1835

M: Serra de Arga: Parque Eólico, larva on Salix atrocinerea Brot., 3-VI-2019, Corley.

Orgyia trigotephras Boisduval, 1829

ALG: Cabranosa, Sagres, larva on Quercus rotundifolia L., 10-IV-2019, Fritsch; E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira.

Callimorpha dominula (Linnaeus, 1758)

DL: Alto das Velhas, 3-VII-2011, Silva.

Coscinia chrysocephala (Hübner, 1804)

BAL: Vargens, Ribeira de Vascão, 24-III-2019, Tagis; R: Colos, 8-VI-2019, Terry.

Lygephila craccae (Denis & Schiffermüller, 1775)

R: Outeiro, Alcobertas, 23-V-2019, Corley.

Tathorhynchus exsiccata (Lederer, 1855)

BL: Lagoa de São José, 26-XII-2019, Rosete.

Parascotia nisseni Turati, 1905

E: Quinta do Pisão, 12-V-2019, Tagis, det. Corley.

Rhypagla lacernaria (Hübner, 1813)

R: Ave Casta, 21-V-2019, Tagis.

Catephia alchymista (Denis & Schiffermüller, 1775)

R: Ave Casta, 21-V-2019, Tagis; E: Lagoa Pequena, Arrimal, 20-VII-2019, Tagis.

Catocala mariana Rambur, 1858

R: Colos, 8-VI-2019, Terry.

Catocala nymphagoga (Esper, 1787)

ALG: Cabranosa, Sagres, larva on Quercus rotundifolia L., 9-IV-2019, Fritsch.

Catocala promissa (Denis & Schiffermüller, 1775)

ALG: Cabo de São Vicente, larva on Quercus rotundifolia L., 10-IV-2019, Fritsch.

Minucia lunaris (Denis & Schiffermüller, 1775)

BAL: Moinhos de Paneiro, 16-IV-2019, Nunes, Silva and Jesus; BA: Faia Brava Reserve, 10-V- 2019, Nunes, Silva and Jesus.

NOCTUIDAE

Chrysodeixis acuta (Walker, 1858)

ALG: Sargaçal, 29-X-2019, Valadares, conf. Yela from photo.

Aedia leucomelas (Linnaeus, 1758)

M: Mosteiro de São João de Arga, 3-VI-2019, Corley and S. Ferreira.

Acronicta megacephala (Denis & Schiffermüller, 1775)

BB: Cortes de Meio, 22-VI-2019, Tagis; BA: Mata do Desterro, 30-VI-2019, Tagis.

Synthimia fixa (Fabricius, 1787)

BAL: Moinhos de Paneiro, 16-IV-2019, Nunes, Silva and Jesus.

Calophasia almoravida Graslin, 1863

BL: Barra de Mira, 3-VI-2019, Tagis.

Omphalophana serrata (Trteitschke, 1835)

BAL: Safara, 12-IV-2019, Tagis.

Lophoterges millierei (Staudinger, 1871)

BAL: Rio Mira at Zambujeira, 29-V-2019, Corley and S. Ferreira; R: Colos, 12-VI-2019, Terry.

Amphipyra tetra (Fabricius, 1787)

TM: Carrazedo, 18-IX-2019, Corley, Nunes and S. Ferreira; São Martinho de Angueira, 23-IX- 2019, Corley and S. Ferreira.

Valeria jaspidea (de Villers, 1789)

ALG: Três Figos, Corsino, 13-III-2019, Tagis; BAL: Santa Bárbara, 11-III-2019, Tagis.

Meganephria bimaculosa (Linnaeus, 1767)

Second to fourth Portuguese records. TM: Carrazedo, 18-IX-2019, Corley, Nunes and S. Ferreira; Guadramil, 20-IX-2019, Corley, Nunes and S. Ferreira; São Martinho de Angueira, 23-IX-2019, Corley and S. Ferreira.

Caradrina selini Boisduval, 1840

ALG: Três Figos, Corsino, 13-III-2019, Tagis.

Caradrina noctivaga Bellier, 1863

R: Colos, 9-VI-2019, Terry.

Hoplodrina octogenaria (Goeze, 1781)

E: Vale da Cabeceira, 25-V-2019, Corley.

Polyphaenis sericata (Esper, 1787)

R: Colos, 8-VI-2019, Terry.

Nonagria typhae (Thunberg, 1784)

TM: Campo de Viboras, 12-VII-2019, Tagis.

Lateroligia ophiogramma (Esper, 1794)

Fourth Portuguese record. DL: Amorim, Póvoa de Varzim, 2-VI-2019, Corley and S. Ferreira.

Mesapamea secalis (Linnaeus, 1758)

BB: Cortes de Meio, 22-VI-2019, Tagis.

Oligia strigilis (Linnaeus, 1758)

BB: Parque Campismo de Pião, 10-VI-2019, Nunes, Silva and Jesus.

Episema grueneri Boisduval, 1837

TM: Estevais, 24-IX-2019, Corley and S. Ferreira.

Spudaea ruticilla (Esper, 1791)

ALG: Cabranosa, Sagres, larva on Quercus rotundifolia L., 14-IV-2019, Fritsch.

Conistra haleae Fibiger & Top-Jenen, 2010

DL: Valongo, 15-II-2019, Nunes.

Scotochrosta pulla (Denis & Schiffermüller, 1775)

TM: São Martinho de Angueira, 23-IX-2019, Corley and S. Ferreira.

Atethmia centrago (Haworth, 1809)

Second Portuguese record. TM: Carrazedo, 18-IX-2019, Corley, Nunes and S. Ferreira.

Polymixis lichenea (Hübner, 1813)

BL: Moura da Serra, 26-X-2019, Rosete.

Lacanobia blenna (Hübner, 1824)

Third Portuguese locality. BL: Quinta do Canal, 1-VI-2019, Corley and Rosete.

Melanchra persicariae (Linnaeus, 1761)

TM: Pitões das Júnias, 24-VII-2019, Nunes and Jesus.

Conisania andalusica (Staudinger, 1859)

R: Valverde, 24-V-2019, Corley and S. Ferreira.

Hadena magnolii (Boisduval, 1829)

BA: Covão da Ametade, 8-VII-2019, Rosete.

Hadena sancta (Staudinger, 1859)

BL: Barra de Mira, 3-VI-2019, Tagis.

Mythimna straminea (Treitschke, 1825)

E: Lagoa de Albufeira, 10-V-2019, Tagis.

Dichagyris forcipula (Denis & Schiffermüller, 1775)

TM: Lamas de Olo, 2-VIII-2018, Gonzalez and Fernandes, det. Corley; Pitões das Júnias, 15-VII- 2019, Nunes and Pires.

* Euxoa nigricans (Linnaeus, 1761)

TM: França, 19-IX-2019, Corley, Nunes and S. Ferreira. Almost all Europe. [Polyphagous on herbaceous plants].

Euxoa obelisca (Denis & Schiffermüller, 1775)

TM: Guadramil, 17-VIII-2019, Nunes, Silva and Jesus.

Axylia putris (Linnaeus, 1761)

E: Vale da Cabeceira, 25-V-2019, Corley.

Noctua fimbriata (Schreber, 1759)

BAL: Lagoa de Santo André, 26-V-2019, Corley.

Noctua tirrenica Biebinger, Speidel & Hanigk, 1983

E: Convento de Arrábida, 30-V-2019, Corley and S. Ferreira.

* Naenia typica (Linnaeus, 1758)

M: Ponte da Cavada Velha, Assureira, Castro Laboreiro, 3-VIII-2018, Jesus and Silva, conf. J. Yela. Widespread in middle latitudes of Europe, but absent from far north and areas with Mediterranean climate. [Polyphagous on herbaceous plants].

NOLIDAE

Nola infantula Kitt, 1926

M: Vale do Cuco, 25-VII-2019, Nunes and Jesus.

Nola tutulella Zerny, 1927

BA: Vilar de Amargo, 11-V-2019, Nunes, Silva and Jesus.

Nycteola columbana (Turner, 1925)

BL: Abiúl, 22-VI-2019, Rosete.

Earias insulana (Boisduval, 1833)

BL: Quinta do Canal, 14-IX-2019, Corley, Rosete, Nunes and S. Ferreira.

Recent literature

BROWN et al. (2019) performed a genetic analysis on a significant number of species of Cochylina (Tortricidae). The main result of this is break up of the polyphyletic genus Cochylis into a number of smaller genera. Details are given below:

Thyraylia Walsingham, 1897

nana (Haworth, 1811)

Cochylis epilinana remains in genus Cochylis for the moment but is not closely related to other Cochylis sensu stricto species (of which there are none in Portugal).

Neocochylis Razowski, 1960

hybridella (Hübner, 1813)

salebrana (Mann, 1862)

dubitana (Hübner, 1799)

molliculana (Zeller, 1847)

Cochylichroa Obraztsov & Swatschek, 1958

atricapitana (Stephens, 1852)

Brevicornutia Razowski, 1960

pallidana (Zeller, 1847)

Pontoturania Obraztsov, 1943

posterana (Zeller, 1847)

CORLEY et al. (2020) describe Mondeguina atlanticella Corley & Rosete, 2020, a new species of Gelechiidae in a new genus from localities near the coast in Beira Litoral and Algarve. Apatetris agenjoi is transferred to genus Catatinagma.

CORLEY & FERREIRA (2019) place Cacochroa rosetella Corley, 2018 in a new genus Rosetea Corley & Ferreira.

CORLEY & FERREIRA (2020) justify the resurrection from synonymy of Cochylimorpha punctiferana (Ragonot, 1881) already proposed in CORLEY (2015).

GAEDIKE (2019) refers Portuguese records of Ceratuncus dzhungaricus to C. maroccanella (Amsel, 1951).

HUEMER (2020) has revised the Caryocolum schleichi species complex. The taxon occurring in Portugal, C. schleichi subspecies dianthella (Chrétien, 1925) is restored to species rank.

LAŠTŮVKA & LAŠTŮVKA (2020) add Bohemannia pulverosella (Stainton, 1849) to the Portuguese fauna.

MARABUTO et al. (2020) study the phylogeography of Euchloe tagis in great detail, relating genetic divergence in this species to refugia in the Western Mediterranean area and geological events over more than five million years. There are no resulting taxonomic changes at species or genus level.

ROSETE et al. (2019) present a list of records from Constância, many of them new records for Ribatejo province.

SLAMKA (2019) makes several nomenclatural changes in the Phycitinae: Neurotomia is synonymised with Melathrix Ragonot, 1893, Moitrelia is synonymised with Uncinus Amsel, 1951 and in genus Elegia the species that has been referred until now as fallax is synonymised with atrifasciella; confusingly fallax is the correct name for fallaximima.

ZLATKOV & HUEMER (2019) raise Clepsis eatoniana (Ragonot, 1881) from synonymy with C. consimilana and demonstrate that it is the oldest name for the recently described C. razowskii Gastón, Vives & Revilla, 2017.

Appendix: Changes to the Portuguese fauna list

Species added to the Portuguese fauna listed in this and other papers are summarised here, each with a number indicating their placement in the checklist (CORLEY, 2015). New genera for the Portuguese fauna show the author and year of publication of the genus.

Name changes resulting from changes at genus level or to new synonymy are given, with each species retaining its list number. In a case where a new name is provided for a previously misidentified species, the new species retains the number of the misidentified species. Thus, Ceratuncus maroccanella (Amsel, 1951) replaces Ceratuncus dzhungaricus Zagulajev, 1971, which was previously misidentified, but the species retains the number 0177 in the checklist.

0063.2 Bohemannia pulverulentella (Stainton, 1849)

0141 Phalacropterix graminifera (Fourcrtoy, 1785) is deleted

0177 Ceratuncus maroccanella (Amsel, 1951) (Ceratuncus dzhungaricus sensu auct. nec Zagulajev, 1971)

0294.1 Phyllonorycter comparella (Duponchel, 1843)

0378.1 Apatema apolausticum Gozmány, 1996

Rosetea Corley & Ferreira, 2019

0579.1 Rosetea rosetella (Corley, 2018) (Cacochroa rosetella Corley, 2018)

Catatinagma Rebel, 1903

0551 Catatinagma agenjoi (Gozmány, 1954) (Apatetris agenjoi Gozmány, 1954)

Mondeguina Corley & Rosete, 2020

0551.1 Mondeguina atlanticella Corley & Rosete, 2020

0626.1 Gladiovalva rumicivorella (Millière, 1881)

0684 Caryocolum dianthella (Chrétien, 1925) (Caryocolum schleichi ssp. dianthella (Chrétien, 1925))

0792.1 Coleophora currucipennella Zeller, 1839

0850.1 Coleophora ribasella Baldizzone, 1982

0897.1 Hypatopa segnella (Zeller, 1873)

SCHRECKENSTEINIOIDEA

SCHRECKENSTEINIIDAE

Schreckensteinia Hübner, 1825

0967.2 Schreckensteinia festaliella (Hübner, 1796)

1006.1 Clepsis eatoniana (Ragonot, 1881) (Clepsis razowskii Gastón, Vives & Revilla, 2017)

1078.1 Aethes mauritanica (Walsingham, 1898)

Thyraylia Walsingham, 1897

1085 Thyraylia nana (Haworth, 1811) (Cochylis nana (Haworth, 1811))

Neocochylis Razowski, 1960

1087 Neocochylis hybridella (Hübner, 1813) (Cochylis hybridella (Hübner, 1813))

1088 Neocochylis salebrana (Mann, 1862) (Cochylis salebrana (Mann, 1862))

1088.1 Neocochylis millierana (Peyerimhoff, 1877)

1089 Neocochylis dubitana (Hübner, 1799) (Cochylis dubitana (Hübner, 1799))

1090 Neocochylis molliculana (Zeller, 1847) (Cochylis molliculana (Zeller, 1847))

Cochylichroa Obraztsov & Swatschek, 1958

1091 Cochylichroa atricapitana (Stephens, 1852) (Cochylis atricapitana (Stephens, 1852))

Brevicornutia Razowski, 1960

1092 Brevicornutia pallidana (Zeller, 1847) (Cochylis pallidana (Zeller, 1847))

Pontoturania Obraztsov, 1943

1093 Pontoturania posterana (Zeller, 1847) (Cochylis posterana (Zeller, 1847))

1166.1 Pseudococcyx friedmariana Larsen, 2020

1480 Elegia atrifasciella Ragonot, 1887 (Elegia fallax sensu auct. nec Staudinger, 1881)

1481 Elegia fallax (Staudinger, 1881) (Elegia fallaximima Nel & Mazel, 2011)

Melathrix Ragonot, 1893

1485 Melathrix coenulentella (Zeller, 1846) (Neurotomia coenulentella (Zeller, 1846))

Uncinus Amsel, 1951

1486 Uncinus obductella (Zeller, 1839) (Moitrelia obductella (Zeller, 1839))

1487 Uncinus hispanella (Staudinger, 1859) (Moitrelia hispanella (Staudinger, 1859))

1662.1 Catoptria lythargyrella (Hübner, 1796)

1921.1 Eupithecia assimilata Doubleday, 1856

2518.1 Euxoa nigricans (Linnaeus, 1761)

Naenia Stephens, 1827

1570.1 Naenia typica (Linnaeus, 1758)

Acknowledgments

We thank all those who have contributed records and Carlos Silva for the photo of Gladiovalva rumicivorella. In particular we thank Patrícia Garcia-Pereira and the Tagis team for the records obtained through the project Invertebrates Red List of Continental Portugal (POSEUR-03-2215-FC- 000094). We are most grateful to Wilfried Arnscheid, José Luis Yela Garcia, Eduardo Marabuto and Martin Townsend for identification or confirmation of specimens and photos, and also to Mike Dale for excellent dissections. Ernestino Maravalhas kindly produced the map. This project has received funding from PORBIOTA - Portuguese E-Infrastructure for Information and Research on Biodiversity (POCI-01-0145-FEDER-022127), supported by Operational Thematic Program for Competitiveness and Internationalization (POCI), under the PORTUGAL 2020 Partnership Agreement, through the European Regional Development Fund (FEDER).

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Author notes

* E-mail: martin.corley@btinternet.com