Artículos

New and interesting Portuguese Lepidoptera records from 2020 (Insecta: Lepidoptera)

Novos e interessantes registos portugueses de Lepidoptera em 2020 (Insecta: Lepidoptera)

Nuevas e interesantes citas portuguesas de Lepidoptera en 2020 (Insecta: Lepidoptera)

Introduction

This paper is the fifteenth in the series of annual summaries of new knowledge of Portuguese Lepidoptera. It gives records of species of Lepidoptera added to the Portuguese fauna in 2020 and some unpublished earlier records, together with new province records not included in the checklist (CORLEY, 2015). Additional data include new data on larval host-plants within the country and second and third records of species for the country, which are only indicated when they are not in new provinces. Papers published in 2020 and part of 2021 that relate to the Portuguese Lepidoptera fauna are listed and briefly summarised. Finally, an Appendix lists the new species for Portugal separately, with numbers indicating their position in the checklist; new genera for Portugal have author and year of publication given.

13 species new for Portugal are listed below, of which three are new for the Iberian Peninsula. Three species are removed from the Portuguese list. A few of the new species listed here have been previously listed for Portugal, but the records were rejected in CORLEY (2015), as being erroneous or unsubstantiated.

In CORLEY et al. (2020) the number of Lepidoptera species recognised from Portugal was 2723. With the current paper and other papers mentioned herein, this total has risen to 2743.

Material and Methods

Most species were captured at light in traps of various kinds, over or beside a white sheet. For specimens not taken at light, the means of capture is given. Specimens are retained in the collections of the original recorders, unless otherwise stated. However, a few records are based only on photographic evidence.

The number of new district records in 2020 is considerably lower than in recent years. Covid 19 restrictions in Portugal and elsewhere and the consequent difficulties of international travel both contributed to a reduction in field work.

The order and nomenclature of families and species follows the Portuguese list (CORLEY, 2015). The nomenclature of plant names follows the EURO+MED PLANT-BASE.

The entry for species new for Portugal concludes with a summary of the known European distribution, and available information on the larval host-plant, given in square brackets if the information comes from outside Portugal.

Localities with UTM squares and altitude: (District in brackets)

List of families and species

NEPTICULIDAE

Stigmella ilicifoliella (Mendes, 1918)

DL: Couce, 22-I-2020, leafminer on Quercus suber L., emerged 3-VII-2020 Nunes, det. Corley.

ADELIDAE

Adela mazzolella (Hübner, 1801)

Second Portuguese record. TM: Ponta da Carba, 9-V-2020, Nunes, det. Corley.

Nematopogon metaxella (Hübner, 1813)

Third Portuguese record. BL: Casal do Redinho, Madriz, 25-V-2020, Rosete.

PSYCHIDAE

Placodoma calpella Sobczyk, 2013

Third Portuguese record. BB: Segura, 10-VI-2020, Silva, Jesus and Nunes, conf. Corley.

Psyche casta (Pallas, 1767)

BL: Louriçal, 18-III-2020, Rosete.

Psyche crassiorella Bruand, 1851

BL: Louriçal, 28-IV-2016, Rosete.

TINEIDAE

** Nemapogon falstriella (Haas, 1881)

BL: Ansião, 20-VI-2020, Rosete. Northern half of Europe, extending south to France, Northern Italy and Croatia. Larvae unknown.

Reisserita zernyi Petersen, 1957

DL: Couce, 3-VII-2020, Nunes, conf. Corley.

Reisserita flavofimbriella (Chrétien, 1925)

BB: Segura, 10-VI-2020, Silva, Jesus and Nunes.

GRACILLARIIDAE

Euspilapteryx auroguttella Stephens, 1825

E: Vale do Figueiredo, 26-V-2020, Cardoso.

Phyllonorycter froelichiella (Zeller, 1839)

BL: Casal do Redinho, Madriz, 25-V-2020, Rosete.

GLYPHIPTERIGIDAE

Orthotelia sparganella (Thunberg, 1788)

BL: Casal do Redinho, Madriz, 25-V-2020, Rosete.

Acrolepiopsis assectella (Zeller, 1839)

E: Pó, 8-X-2020, Cardoso; M: Senhora da Madalena, Lindoso, 17-VII-2020, Silva, Jesus and Nunes.

LYONETIIDAE

Leucoptera andalusica Mey, 1994

DL: Couce, larvae mining cladodes of Genista tridentata L., 25-I-2020, Nunes.

PRAYDIDAE

Prays citri (Millière, 1873)

E: Pó, 5-V-2020, Cardoso.

AUTOSTICHIDAE

Oegoconia quadripuncta (Haworth, 1828)

DL: Areia, 14-V-2020, Jesus, det. Corley.

Symmoca alhambrella Walsingham, 1911

BL: Casmilo, 10-VII-2020, Rosete.

LECITHOCERIDAE

Homaloxestis briantiella (Turati, 1879)

M: Senhora da Madalena, Lindoso, 17-VII-2020, Silva, Jesus and Nunes, det. Corley.

OECOPHORIDAE

Batia lambdella (Donovan, 1793)

E: Pó, 2-VI-2020, Cardoso.

* Metalampra italica Baldizzone, 1977

DL: Areia, 14-V-2020, Jesus, conf. Corley. Described from Italy in 1977 but has recently spread to Croatia and northern and western parts of Europe, reaching Germany, Netherlands, England, Wales, France and Spain (GASTÓN & VIVES MORENO, 2020: 310). [Larva in fungus-infected decaying wood].

Pleurota andalusica Back, 1973

DL: Couce, 3-VII-2020, Nunes, det. Corley.

DEPRESSARIIDAE

Agonopterix ocellana (Fabricius, 1775)

TM: Ponte da Carba, 8-V-2020, Silva, Jesus and Nunes.

Agonopterix olusatri Corley & Buchner, 2019

BB: Segura, 10-VI-2020, Silva, Jesus and Nunes.

Depressaria depressana (Fabricius, 1775)

BL: Castelo do Germanelo, 5-IX-2020, Rosete.

Odites kollarella (Costa, 1832)

M: Senhora da Madalena, Lindoso, 17-VII-2020, Silva, Jesus and Nunes.

COSMOPTERIGIDAE

Anatrachyntis badia (Hodges, 1962)

E: Pó, 12-V-2020, Cardoso.

Coccidiphila gerasimovi Danilevsky, 1950

DL: Águas Santas, Maia, 14-III-2020, Nunes, det. Corley.

Eteobalea dohrnii (Zeller, 1847)

BL: Abiúl, 15-VII-2020, Rosete.

GELECHIIDAE

Stomopteryx basalis Staudinger, 1876

BL: Poço, 23-VII-2020, Rosete.

Aproaerema buvati (Nel, 1995)

BL: Abiúl, larva on Coronilla repanda (Poir.) Guss., 31-V-2019, Corley and Rosete.

Aproaerema polychromella (Rebel, 1902)

BB: Segura, 10-VI-2020, Silva, Jesus and Nunes; M: Senhora da Madalena, Lindoso, 17-VII-2020, Silva, Jesus and Nunes.

Anacampsis timidella (Wocke, 1887)

DL: Couce, 3-VII-2020, Nunes, det. Corley.

Mondeguina atlanticella Corley & Rosete, 2020

Extension of range. BL: Cais do Moliço, Vagos, 18-VII-2020, Rosete.

Aristotelia decoratella (Staudinger, 1879)

BL: Ansião, 7-VIII-2020, Rosete.

Pyncostola bohemiella (Nickerl, 1871)

Second and third Portuguese records. BL: Abiúl, 15-VII-2020, Rosete; Ansião, 7-VIII-2020, Rosete.

Neofriseria peliella (Treitschke, 1835)

M: Travanca, 18-VII-2020, Silva. Jesus and Nunes, det. Corley.

Mirificarma cytisella (Treitschke, 1833)

BB: Segura, 10-VI-2020, Silva. Jesus and Nunes.

Mirificarma interrupta (Curtis, 1827)

BL: Louriçal, flying at dusk, 9-IV-2020, Rosete.

Caryocolum fibigerium Huemer, 1988

M: Travanca, 18-VII-2020, Silva. Jesus and Nunes, det. Corley.

Teleiodes albidorsella Huemer & Karsholt, 1999

DL: Couce, 3-VII-2020, Nunes, det. Corley.

Schistophila laurocistella Chrétien, 1899

TM: Ribeira Angueira, 9-V-2020, Nunes, det, Corley.

ELACHISTIDAE

Elachista boursini Amsel, 1951

BB: Segura, 10-VI-2020, Silva, Jesus and Nunes, det. Corley.

Elachista igaloensis Amsel, 1951

DL: Couce, leafmine on Carex sp., 8-I-2020, Nunes, det. Kaila (fig. 1).

** Elachista differens Parenti, 1978

BL: Pampilhosa de Botão, 17-V-2018, Jesus; Abiúl, 2-IX-2019, Rosete, det. Corley. France, Italy, Switzerland, Germany. [Leaf-miner on Carex ferruginea Scop.].

COLEOPHORIDAE

Coleophora trifariella Zeller, 1849

Records from Beira Litoral (PIRES & CORLEY, 2007) and from Minho (CORLEY et al., 2016) are misidentifications of Coleophora genistae Stainton, 1857. Mendes’ records from Beira Baixa are correct.

Coleophora sisteronica Toll, 1961

E: Salir do Porto, reared from Coronilla valentina L., 7-VII-2020, Nunes, conf. Corley.

Coleophora flaviella Mann, 1857

TM: Ribeira Angueira, 9-V-2020, Nunes, det, Corley.

Coleophora galbulipennella Zeller, 1838

Fourth Portuguese record. TM: Montesinho, 24-VIII-2020, Jesus, Fernandes and Nunes, det. Corley.

SCYTHRIDIDAE

** Scythris sinensis (Felder & Rogenhofer, 1875)

M: Adaúfe, 10-VIII-2020, Marques, det. Nunes from photo on Facebook. China, Japan, Korea and Siberia; in Eastern Europe known from Russia, Estonia, Latvia, Lithuania, Belorussia, Ukraine, Moldova, Hungary and Poland, apparently increasing its range. Records from England and Italy are probably accidental imports. [Chenopodium album L., Atriplex patula L.].

Scythris subseliniella (Heinemann, 1876)

M: Travanca, 18-VII-2020, Silva, Jesus and Nunes, det. Corley

Enolmis acanthella (Godart, 1825)

E: Pó, 27-IV-2020, Cardoso.

BLASTOBASIDAE

Hypatopa segnella (Zeller, 1873)

BA: Picão, 1-VIII-2020, Silva, Jesus and Nunes, det. Corley; M: Senhora da Madalena, Lindoso, 17-VII-2020, Silva, Jesus and Nunes.

MOMPHIDAE

Mompha propinquella (Stainton, 1851)

BL: Porta da Sula, 30-VII-2020, Jesus, det. Nunes.

ALUCITIDAE

* Alucita grammodactyla Zeller, 1841

ALG: Praia da Rocha, 3-III-2020, Hellers, det. Corley. South and middle latitudes of Europe north to south Sweden; absent from Mediterranean islands, Greece and British Isles. [Scabiosa columbaria L., S. canescens Waldst. & Kit.].

SCHRECKENSTEINIIDAE

Schreckensteinia festaliella (Hübner, 1796)

DL: Bajouca, Gemunde, 16-VII-2020, Silva.

EPERMENIIDAE

Epermenia aequidentellus (E. Hofmann, 1867)

TM: Santuário de Santo Ambrósio, larvae on Pimpinella villosa Schousb., 10-V-2020, Nunes, det. Corley

TORTRICIDAE

Cnephasia stephensiana (Doubleday, 1849)

TM: Olmares, reared from Andryala integrifolia L., 31-V-2020, Nunes, det. Corley

Acleris lorquiniana (Duponchel, 1843)

Seond Portuguese locality. BL: Casal do Redinho, Madriz, 25-V-2020, Rosete.

Aethes francillana (Fabricius, 1794)

ALG: Carrapateira, emerged 5-IX-2019 from seeds of Crithmum maritimum L., collected 21-VIII2019, Rosete, det. Corley

Apotomis betuletana (Haworth, 1811)

Third Portuguese record. TM: Montesinho, 24-VIII-2020, Nunes, Jesus and Fernandes.

Lobesia artemisiana (Zeller, 1847)

AAL: Ramila, 3-VII-2020, Valadares.

Gypsonoma imparana Müller-Rutz, 1914

TM: Vilarinho das Azenhas, 25-IX-2020, Nunes, Ferreira and Mata, det. Corley.

* Epiblema foenella (Linnaeus, 1758)

BA: Ribeira da Carvalhosa, Picão, 7-VII-2020, Teixeira (fig. 2). Nearly all Europe except Mediterranean islands and Ireland. [Larva in rootstock of Artemisia vulgaris L., A. abrotanum L. and Cota tinctoria (L.) J. Gay].

Pseudococcyx tessulatana (Staudinger, 1871)

BL: Castelo do Germanelo, 29-V-2020, Rosete.

Dichrorampha chavanneana (La Harpe, 1858)

DL: Quintarei, 13-III-2020, Nunes, det. Corley

Grapholita janthinana (Duponchel, 1843)

M: Senhora da Madalena, Lindoso, 17-VII-2020, Silva, Jesus and Nunes

HETEROGYNIDAE

Delete Heterogynis penella (Hübner, 1819) from the Portuguese list. There is no evidence that this species occurs in Portugal. Most probably all records belong to H. cynetis de Freina et al., 2020.

Heterogynis cynetis de Freina, Monasterio, Escobés, Hinojosa & Vila, 2020

TM: Balsamão, 9-VI-2016, Nunes.

PYRALIDAE

Achroia grisella (Fabricius, 1794)

ALG: Sargaçal, 11-VIII-2017, Valadares, det. Nunes.

Lamoria anella (Denis & Schiffermüller, 1775)

M: Senhora da Madalena, Lindoso, 17-VII-2020, Silva, Jesus and Nunes.

Pyralis regalis (Denis & Schiffermüller, 1775)

BA: Poço do Inferno, 16-VIII-2020, Rosete, conf. Corley.

Stemmatophora vulpecalis (Ragonot, 1891)

BB: Segura, 10-VI-2020, Silva, Jesus and Nunes.

* Valdovecaria hispanicella (Herrich-Schäffer, 1855)

AAL: Hortas de Baixo, 20-IX-1995, Corley; Ramila, 1-VII-2020, Valadares, det. Marabuto from photo (fig. 3). Spain, France. Larva unknown.

Asalebria florella (Mann, 1862)

M: Senhora da Madalena, Lindoso, 17-VII-2020, Silva, Jesus and Nunes.

Elegia fallax (Staudinger, 1881) sensu Slamka, 2019

TM: Ponte da Carba, 8-V-2020, Nunes, det. Corley.

Acrobasis fallouella (Ragonot, 1871)

Third Portuguese record. BA: Picão, 1-VIII-2020, Silva, Jesus and Nunes, det. Corley.

Acrobasis glaucella Staudinger, 1859

DL: Valongo, reared from Quercus suber L., 17-VI-2020, Nunes, det. Corley.

Euzophera osseatella (Treitschke, 1832)

E: Pó, 21-V-2020, Cardoso.

Delete Euzopherodes vapidella (Mann, 1857) from the Portuguese list. NEL et al., (2021) describe a new species Euzopherodes vapidelloides from France including Corsica and from Spain, distinct from E. vapidella. We have examined Portuguese specimens of Euzopherodes and found them to belong to the new species. There is no evidence that E. vapidella occurs in Portugal and it should be deleted from the Portuguese list. E. vapidella is known from Madrid, so its presence in Portugal remains a possibility.

* Euzopherodes vapidelloides Labonne, Nel & Varenne, 2021

Portuguese specimens previously identified as E. vapidella (Mann, 1857), with records from ALG, AAL, BB and TM, belong to this newly described species. Spain, France, Corsica. Larva unknown.

Phycitodes albatella (Ragonot, 1887)

M: Louredo, 17-X-2020, Nunes, det. Corley.

Ephestia disparella Hampson, 1901

M: Travanca, 18-VII-2020, Silva. Jesus and Nunes.

CRAMBIDAE

Metasia ibericalis Ragonot, 1894

BAL: Herdade da Varginha, 28-VI-2020, Jesus.

Metasia cuencalis Ragonot, 1894

M: Senhora da Madalena, Lindoso, 17-VII-2020, Silva, Jesus and Nunes.

Mesocrambus tamsi Bleszynski, 1960

BAL: Herdade da Varginha, 28-VI-2020, Jesus, conf. Nunes.

* Mesocrambus marabut ( Bleszynski, 1965)

BL: Montes Santa Olaia, 30-VII-2003, Pires; ALG: Malhada da Figueira, 1-VIII-2011, Montegriffo, det. Corley. This species has been misdentified as M. carectellus since MONTEIRO (1982). It is almost certain that all records of M. caractellus belong here. Spain. Larva unknown.

Delete Mesocrambus carectellus (Zeller, 1847) from the Portuguese list, in which it was included by MONTEIRO (1982), but recent re-examination of a number of specimens suggests that all records belonging to M. marabut.

DREPANIDAE

* Ochropacha duplaris (Linnaeus, 1761)

M: Castro Laboreiro, 9-VII-2019, Valadares (fig.4). Nearly all Europe, but absent from Mediterranean islands and some Balkan countries. [Alnus, Betula, Corylus, Populus, Quercus].

LASIOCAMPIDAE

Trichiura castiliana Spuler, 1908

ALG: Sargaçal, 8-XI-2014, Valadares.

GEOMETRIDAE

Idaea circuitaria (Hübner, 1819)

In CORLEY (2015) the first record for Portugal is given from Setúbal, but E was accidentally omitted from the list of provinces.

Cyclophora quercimontaria (Bastelberger, 1897)

M: Fradelos, 24-IV-2020, Nunes, det. Corley

Cyclophora linearia (Hübner, 1799)

BA: Picão, 1-VIII-2020, Silva, Jesus and Nunes, det. Corley; DL: Bouças de Arões, Bajouca, Gemunde, 30-V-2020, Silva, Jesus, Nunes and Ferreira.

Anticlea derivata (Denis & Schiffermüller, 1775)

TM: São Joanico, 21-III-2019, Cordeiro.

* Minoa murinata (Scopoli, 1763)

TM: Lama Grande, 20-V-2020, Teixeira (fig. 5). Most of Europe, but absent from Scandinavia, Baltic states, Ireland, Balearic Islands, Sardinia and Crete. [Euphorbia species, particularly E. cyparissias L. and E. amygdaloides L.].

Lobophora halterata (Hufnagel, 1767)

Third Portuguese record. TM: Ponte da Carba, 21-V-2020, Teixeira.

Isturgia deerraria (Walker, 1861)

BL: Pampilhosa de Botão, 16-IX-2020, Jesus.

Agriopis marginaria (Fabricius, 1776)

E: Vale de Figueiredo, Camarnais, 23-I-2020, Cardoso.

Lomographa bimaculata (Fabricius, 1775)

TM: Gontães, 23-V-20, Fernandes and Gonzalez, det. Nunes.

Ortaliella perspersata (Treitschke, 1827)

BB: Segura, 10-VI-2020, Silva, Jesus and Nunes.

Phaiogramma etruscaria (Zeller, 1849)

BAL: Herdade da Varginha, 28-VI-2020, Jesus.

NOTODONTIDAE

Stauropus fagi (Linnaeus, 1758)

ALG: Alfambras, 13-VI-2020, Valadares.

EREBIDAE

Gynaephora fascelina (Linnaeus, 1758)

Second Portuguese record. TM: Lama Grande, 16-VII-2020, Teixeira.

Callimorpha dominula (Linnaeus, 1758)

TM: Nogueira, 20-V-2017, Fernandes and Gonzalez, det. Nunes.

Coscinia striata (Linnaeus, 1758)

E: Planalto das Cortes, Camarnais, 19-V-2020, Cardoso.

Coscinia chrysocephala (Hübner, 1804)

E: Vale de Figueiredo, Camarnais, 21-VI-2016, Cardoso.

Catocala nupta (Linnaeus, 1767)

ALG: Foz do Ribeiro, 3-VII-2016, Valadares.

NOCTUIDAE

Plusia festucae (Linnaeus, 1758)

ALG: Sargaçal, 5-VI-2015, Valadares.

* Xanthodes albago (Fabricius, 1794)

ALG: Sargaçal, 16-VI-2020, Valadares (fig. 6|). Mediterranean species, mainly near the coast from Spain to Greece. [Malva olbia (L.) Alef., M. moschata L.].

Acronicta alni (Linnaeus, 1767)

TM: Ponte do Couço, 16-V-2020, Silva, Jesus and Nunes.

Bryophila vandalusiae Duponchel, 1842

BB: Sipote, Ermida, 5-IX-2020, Ramos, conf. Corley.

Agrochola helvola (Linnaeus, 1758)

TM: Ponta da Carba, larva on Quercus pyrenaica Willd., 9-V-2020, emerged 17-X-2020, Nunes, conf. Corley.

Agrochola lota (Clerck, 1759)

E: Vale de Figueiredo, Camarnais, 1-I-2020, Cardoso.

Polymixis lichenea (Hübner, 1813)

E: Cesaredas, Reguengo Grande, 13-XI-2020, Cardoso.

Hadena magnolii (Boisduval, 1829)

E: Pó, 29-IV-2020, Cardoso.

* Mythimna pallens (Linnaeus, 1758)

BL: Casais do Porto, 13-VI-2020, Rosete (figs 7, 8) Almost all Europe including Sicily, but absent from other Mediterranean islands. [Grasses].

Mythimna languida (Walker, 1858)

BAL: Herdade da Varginha, 28-VI-2020, Jesus.

Recent literature

ARNSCHEID & WEIDLICH (2021) review the taxonomy of the Ptilocephala albida-species group in Europe, concluding that there are three species. P. albida sensu stricto is found in France, south-west Germany, Switzerland and northern Italy, but not in the Iberian Peninsula. In Portugal and Spain the group is represented by P. lorquiniella (Bruand, 1853).

CORLEY & FERREIRA (2021) describe Ypsolopha milfontensis from the Alentejo coast, a new species with Ephedra fragilis Desf. as host-plant.

CORLEY et al. (2020a) raise Agonopterix rigidella (Chrétien, 1907) from synonymy with A. fruticosella. Both species occur in Portugal: A. fruticosella is recorded from Algarve; A. rigidella is recorded from Algarve and Estremadura.

CORLEY et al. (2020b) add 14 species to the Portuguese list and delete two.

CORRO CHANG & METZ (2021) transfer Tuta absoluta to genus Phthorimaea, based on a cladistic analysis of Gnorimoschemini.

FREINA et al. (2020) describe Heterogynis cynetis from Algarve. This is wholly or in part the species referred to in CORLEY (2015) as H. penella (Hübner, 1819). It has still to be ascertained whether Heterogynis from lowland localities further north in Portugal belong to the new species or another and if there is only one species in Algarve, but there is no reason to suppose that H. penella occurs in Portugal, and it should therefore be removed from the Portuguese list.

HUEMER & KARSHOLT (2020) place Monochroa melagonella in the genus Pragmatodes Walsingham, 1908. Monochroa nigromaculella (Millière, 1872) is transferred to Oxypteryx Rebel, 1911.

MOYA et al. (2017) made a very thorough genetic investigation of the relationships of Argynnis Fabricius and Speyeria Scudder. Their recommendation was that three genera should be recognised, Argynnis, Fabriciana and Speyeria but the justification for this recommendation was a reluctance to alter the names of numerous Speyeria species in North America to Argynnis. There were three lineages found, but Fabriciana and Speyeria are nested within Argynnis, so recognition of three genera is not justified on evolutionary grounds. For the Portuguese species, following their recommendation would mean changing the names of three of the five species. We favour retention of the name Argynnis for all five species, but suggest that the genera they recognise should be treated as subgenera.

NUNES (2021) adds new information on host-plants of 100 species of Lepidoptera within Portugal.

NUNES et al. (2021) add six new species for Portugal: Aplota palpellus (Haworth, 1828), Anacampsis obscurella (Denis & Schiffermüller, 1775), Coleophora anitella Baldizzone, 1985, Apomyelois bistriatella (Hulst, 1887), Eublemma amoena (Hübner, 1803) and Xestia sexstrigata (Haworth, 1809).

WIKSTRÖM et al. (2020) revise the Pyralis regalis group, raising subsp. sagarrai Leraut, 2005 to species rank and adding it to the Portuguese fauna, with records from Alto Alentejo and Trás-os-Montes.

Acknowledgments

Special thanks to Helder Cardoso, Ana Valadares and Edmundo Jesus for the contribution of numerous records. We are also most grateful to all those others who have contributed records; to Lauri Kaila and Eduardo Marabuto who identified or confirmed specimens and photos; to Mike Dale for excellent dissections and to Ernestino Maravalhas who kindly produced the map.

BIBLIOGRAPHY

ARNSCHEID, W. R. & WEIDLICH, M., 2021.– First taxonomic revision of the Ptilocephala albida-species group in Europe (Lepidoptera: Psychidae, Oiketicinae).– SHILAP Revista de lepidopterología, 49(195): 417-433.

CORLEY, M. F. V., BUCHNER, P., RYMARCZYK, F. & DUTHEIL, M., 2020a.– Agonopterix rigidella (Chrétien, 1907) resurrected from synonymy (Lepidoptera, Depressariidae).– Entomologist’s Record and Journal of Variation, 132: 281-296.

CORLEY, M. & FERREIRA. S., 2021.– Ypsolopha milfontensis, a new species from the Portuguese coast (Lepidoptera, Ypsolophidae).– SHILAP Revista de lepidopterología, 49(193): 55-64.

CORLEY, M. F. V., NUNES, J., ROSETE, J., TERRY, R. & FERREIRA, S., 2020b.– New and interesting Portuguese Lepidoptera records from 2019 (Insecta: Lepidoptera).– SHILAP Revista de lepidopterología, 48(192): 609-641.

CORLEY, M. F. V., ROSETE, J., GONÇALVES, A. R., NUNES, J., PIRES, P. & MARABUTO, E., 2016.– New and interesting Portuguese Lepidoptera records from 2015 (Insecta: Lepidoptera).– SHILAP Revista de lepidopterología, 44(176): 615-643.

CORRO CHANG, P. & METZ, M., 2021.– Classification of Tuta absoluta (Meyrick, 1917) (Lepidoptera: Gelechiidae: Gelechiinae: Gnorimoschemini) based on cladistic analysis of morphology.– Proceedings of the Entomological Society of Washington, 123(1): 41-54.

FREINA, J. de, MONASTERIO-LÉON, Y., ESCOBÉS-JIMÉNEZ, R., HINOJOSA, J. C. & VILA, R., 2020.– Heterogynis Rambur, 1837 (Lepidoptera: Zygaenoidea, Heterogynidae) from Andalusia and the Algarve (Spain and Portugal): a taxonomic and faunistic review.– Boletín de la Sociedad Entomológica Aragonesa, 67: 75-85.

GASTÓN, J. & VIVES MORENO, A., 2020.– Contribución al conocimiento de los Lepidoptera de España con la descripción de cuatro nuevas especies para nuestra fauna y otras citas de interés (Insecta: Lepidoptera).– SHILAP Revista de lepidopterología, 48(190): 307-324.

HUEMER, P. & KARSHOLT, O., 2020.– Commented Checklist of European Gelechiidae (Lepidoptera).– ZooKeys, 921: 65-140.

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Notas de autor

*Autor para la correspondencia / Corresponding author martin.corley@btinternet.com